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1.
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Apical clypeal angle with 1 small, sharp, clearly upturned tooth on each side of apical emargination (Fig. 1) |
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1'.
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Clypeus rounded or angled at each side of median emargination, always lacking teeth (Fig. 2) |
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Figure 1. Odontopsammodius cruentus. |
Figure 2. Tesarius caelatus.
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2(1). |
Dorsal surface lacking setae |
3 |
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2'.
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Dorsal surface with setae throughout (Fig. 3) |
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Figure 3. Trichopsammodius brasiliensis. |
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3(2).
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Body robust, somewhat globose (Fig. 4). Basal metatarsomere short, triangularly expanded |
Odontopsammodius Gordon and Pittino |
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3'.
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Body elongate, somewhat cylindrical (Fig. 5). Basal metatarsomere weakly shortened, widened apically, usually not notably triangular |
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Figure 4. Odontopsammodius cruentus. |
Figure 5. Parapsammodius pseudointeger. |
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4(2).
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Pronotal punctures large, simple, with depressed setal pore. Elytral setae prominent (Fig. 6). Metafemur broad, wider than profemur. Metatarsomeres strongly, obviously asymmetrically widened apically |
Trichiopsammodius Petrovitz |
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4'.
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Pronotal punctures very large, appearing O-shaped, with small central tubercle bearing setae. Elytral setae short, indistinct (Fig. 7). Metafemur slender, obviously narrower than profemur. Metatarsomeres moderately, asymmetrically widened apically |
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Figure 6. Trichopsammodius brasiliensis. |
Figure 7. Mysarus peruanus. |
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5(1). |
Metatibiae with distinct transverse carina (Fig. 8) |
Tesarius Rakovic |
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5'.
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Metatibia lacking transverse carina, rarely with weak transverse line, often with row of setiferous teeth (Fig. 9) |
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Figure 8. Tesarius caelatus. |
Figure 9. Geopsammodius rileyi. |
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6(5).
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Pronotal sculpture with at least 5 transverse ridges (Fig. 10), ridges either continuous or broken into discrete granules, usually with as many transverse furrows |
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7 |
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6'.
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Pronotal sculpture with at most 3 weak, rarely with vague 4, transverse ridges and furrows (Fig. 11) |
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Figure 10. Psammodius basalis. |
Figure 11. Leiopsammodius malkini. |
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7(6).
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Metatarsomeres with at least basal segment short, broad, strongly asymmetrically widened apically (Fig. 12) |
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8 |
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7'.
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Metatarsomeres elongate, basal segment subcylindrical (as in Fig. 13), slightly symmetrically or asymmetrically widened apically |
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Figure 12. Psammodius basalis. |
Figure 13. Pleurophorus caesus. |
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8(7).
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Vertex of head with 1 or 2 pairs of oblique ridges arranged in chevron (Fig. 14), ridges either continuous or broken into discrete tubercles |
...
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Psammodius Fallen |
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8'.
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Vertex of head without oblique ridges (Fig. 15)
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Figure 14. Psammodius basalis. |
Figure 15. Neopsammodius werneri. |
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9(7).
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Elytra with scale-like setae on raised intervals (Fig. 16) |
Trichiorhyssemus Clouët |
Figure 16. Trichiorhyssemus riparus. |
9'.
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Elytra lacking setae |
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10(9).
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Vertex of head with 1 or 2 pairs of oblique ridges arranged in a chevron, ridges either continuous or broken into discrete tubercles (Fig. 17). Pronotal margins obviously setaeous. Elytral intervals either flattened or costate with distinct rows of fine to coarse tubercles |
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Rhyssemus Mulsant |
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10'.
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Vertex of head without oblique ridges (Fig. 18). Pronotal margins lacking setae. Odd elytral intervals strongly costate, even intervals flat, both lacking tubercles |
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Figure 17. Rhyssemus germanus. |
Figure 18. Neorhyssemus quinquecostatus. |
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11(6).
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Body short, robust, visibly dilated posteriorly (Fig. 19). Elytral base not margined. Posterior tarsus short, tarsomeres 1-4 triangular (Fig. 20). |
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12 |
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11'.
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Body slender, elongate, either suboval or subparallel, slightly dilated posteriorly (Fig. 21). Elytral base margined. Posterior tarsus elongate or somewhat shortened, all tarsomeres elongate or with basal 1 or 2 segments asymmetrically widened apically (Fig. 22) |
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Figure 19. Leiopsammodius malkini. |
Figure 20. Leiopsammodius malkini. |
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Figure 21. Platytomus micros. |
Figure 22. Platytomus micros. |
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12(11).
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Eye normal. Frontal suture usually distinct (Fig. 23). Tenth elytral interval reaching apical third (Fig. 24). Wing normally developed. Claw corneous |
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Leiopsammodius Rakovic |
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12'.
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Eye small, poorly developed. Frontal suture apparently lacking (Fig. 25), vertex lacking transverse ridge. Tenth elytral interval short, only attaining middle of eltyra (Fig. 26). Wing vestigial. Claw setaceous |
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Figure 23. Leiopsammodius acei. |
Figure 24. Leiopsammodius deyrupi. |
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Figure 25. Geopsammodius hydropicus. |
Figure 26. Geopsammodius morrisi. |
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13(11).
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All elytral intervals ridged to costate (Fig. 27). Pronotum with 3 more or less distinct, flat, vestigial transverse ridges, with equal number of transverse furrows, distinct vestige of midline furrow present |
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Afrodiasticus Pittino and Mariani |
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13'.
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All elytral intervals weekly convex to flat, never ridged to costate (Fig. 28). Pronotum with no transverse ridges, with at most 2 incomplete transverse furrows, vestige of midline furrow present |
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Figure 27. Afrodiastictus mimicus. |
Figure 28. Platytomus micros. |
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14(13).
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Metatarsus elongate, at least as long as tibia; first tarsomere subequal in length to long spur (Fig. 29), cylindrical, slightly widened apically |
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Pleurophorus Mulsant |
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14'.
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Metatarsus weakly to strongly shortened, visibly shorter than tibia; first tarsomere shorter than long spur (Fig. 30), weakly to visibly widened at apex and slightly asymmetrical |
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Figure 29. Pleurophorus caesus. |
Figure 30. Platytomus micros. |
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