Type genus: Anaides Westwood, 1845.

Description. Length 2.0-14.0 mm. Shape variable. Color brown, black, or reddish brown. Head not deflexed. Antennae 10-segmented with 3-segmented, opposable club; club oval, first segment cupuliform or not. Eye canthus variably developed. Eyes developed. Labrum produced beyond apex of clypeus, prominent. Mandibles produced beyond apex of labrum, thick at base, apex pointed, external surface generally sculptured. Maxillae with 4-segmented palpi; labium with apex indented, with 4-segmented palpi (rarely 3-segmented). Pronotum convex or flat, generally sculptured or punctate; base sinuous or with triangular projection. Scutellum exposed, triangular. Elytra convex or flat, generally sculptured, striae well-defined. Pygidium concealed by elytra. Hind wings well-developed or reduced, with M-Cu loop present or absent, MP3 vein present, and MP4 vein present or absent. Venter with sculpturing strigulate or vermiculate. Prosternum bi-concave. Mesosternum not invaginated. Abdomen with 5-6 free sternites (first sternites obscured by hind coxae except lateral edges) and 8 functional spiracles; spiracles 1-7 situated on pleural membrane, spiracle 8 situated in tergite. Protibiae with 2 or 3 teeth. Meso- and metatibia without transverse carinae. Tarsi 5-5-5. Ventral position of protarsi at or apical of second tibial tooth. Tarsal claws simple. Male genitalia symmetrical (rarely slightly asymmetrical).

Composition. The subfamily Anaidinae includes seven genera (one fossil) and 59 species (two fossil).


Distribution. Neotropical; subregions Caribbean, Amazonica, and Chaqueña. The fossil genus Cretanaides Nikolajev is from the Cretaceous of Siberia.

Natural history. Adults feed primarily on dung and carrion, and some species feed on fungi. Most species are attracted to lights. Larvae have been collected under bark and in decomposed plant material.

Ocampo, F. C. 2006. Phylogenetic Analysis of the Scarab Family Hybosoridae and Monographic Revision of the New World Subfamily Anaidinae (Coleoptera: Scarabaeidae). Bulletin of the University of Nebraska State Museum, Vol. 19.