Type
genus: Anaides Westwood, 1845.
Description. Length 2.0-14.0 mm. Shape
variable. Color brown, black, or reddish brown. Head not deflexed. Antennae 10-segmented
with 3-segmented, opposable club; club oval, first segment cupuliform or not.
Eye canthus variably developed. Eyes developed. Labrum produced beyond apex of
clypeus, prominent. Mandibles produced beyond apex of labrum, thick at base, apex
pointed, external surface generally sculptured. Maxillae with 4-segmented palpi;
labium with apex indented, with 4-segmented palpi (rarely 3-segmented). Pronotum
convex or flat, generally sculptured or punctate; base sinuous or with triangular
projection. Scutellum exposed, triangular. Elytra convex or flat, generally sculptured,
striae well-defined. Pygidium concealed by elytra. Hind wings well-developed or
reduced, with M-Cu loop present or absent, MP3 vein present, and MP4 vein present
or absent. Venter with sculpturing strigulate or vermiculate. Prosternum bi-concave.
Mesosternum not invaginated. Abdomen with 5-6 free sternites (first sternites
obscured by hind coxae except lateral edges) and 8 functional spiracles; spiracles
1-7 situated on pleural membrane, spiracle 8 situated in tergite. Protibiae with
2 or 3 teeth. Meso- and metatibia without transverse carinae. Tarsi 5-5-5. Ventral
position of protarsi at or apical of second tibial tooth. Tarsal claws simple.
Male genitalia symmetrical (rarely slightly asymmetrical).
Composition. The subfamily Anaidinae
includes seven genera (one fossil) and 59 species (two fossil).
Genera
Anaides
Callosides
Chaetodus
Cryptogenius
Hybochaetodus
Totoia
Distribution. Neotropical; subregions
Caribbean, Amazonica, and Chaqueña. The fossil genus Cretanaides Nikolajev
is from the Cretaceous of Siberia.
Natural history. Adults feed primarily
on dung and carrion, and some species feed on fungi. Most species are attracted
to lights. Larvae have been collected under bark and in decomposed plant material.
Bibliography
Ocampo, F. C. 2006. Phylogenetic Analysis of the Scarab Family Hybosoridae and
Monographic Revision of the New World Subfamily Anaidinae (Coleoptera: Scarabaeidae).
Bulletin of the University of Nebraska State Museum, Vol. 19. 210pp.