TAXA
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PLEOCOMIDAE
PASSALIDAE
GEOTRUPIDAE
BELOHINIDAE
GLARESIDAE
TROGIDAE
DIPHYLLOSTOMATIDAE
LUCANIDAE
HYBOSORIDAE
OCHODAEIDAE
GLAPHYRIDAE

Aphodiinae
Scarabaeinae
Pachypodinae
Orphinae
Allidiostomatinae
Dynamopodinae
Aclopinae
Euchirinae
Phaenomeridinae
Melolonthinae
Rutelinae
Dynastinae
Cetoniinae