Henry F. Howden 1925-2014
 
howden.jpg

Henry Howden at the national meetings
of the Entomological Society of America,
Detroit, Michigan, December 1983.
Photo by Art Evans.



  

Henry Howden had broad interests in the taxonomy, ecology, biology, biogeography, and evolution of the Scarabaeoidea. During his long career he published numerous papers on many groups of scarabs, and he was a dedicated field biologist. He was probably best known as the world's leading specialist on the Geotrupinae, and he published major monographs on the geotrupines of North, Central, and South America as well as Australia. He also conducted extensive taxonomic research on the Scarabaeinae and produced a monograph (with O. Young) on the Scarabaeinae of Panama and a monograph (with Oscar Cartwright) on the Onthophagus of the United States. Henry retired from Carlton University and remained in Ottawa for several years before moving to Victoria, British Columbia, Canada. His extensive collection was donated to the Canadian Museum of Nature. His passing marks the end of an era in scarabaeoid research.

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  PUBLICATIONS
 
  1. Howden, H.F. 1946. The brown king snake or mole snake (Lampropeltis rhombomaculata) in Maryland. Maryland, A Journal of Natural History 16: 38-40.
  2. Howden, H.F., A.T. Howden, and P.O. Ritcher. 1951. Insects feeding on poison oak (Rhus toxicodendron L.). The Coleopterists Bulletin 5: 17-19.
  3. Howden, H.F. and G.B. Vogt. 1951. Insect communities of standing dead pine (Pinus virginiana Mill.). Annals of the Entomological Society of America 44: 581-595.
  4. Howden, H.F. and R. Mansueti. 1951. Fishes of the tributaries of the Anacostia River, Maryland. Proceedings of the Biological Society of Washington 64: 93-96.
  5. Howden, H.F. 1952. A new name of Geotrupes (Peltotrupes) chalybaeus LeConte, with a description of the larva and its biology (Scarabaeidae). The Coleopterists Bulletin 6: 41-48.
  6. Howden, H.F. and P.O. Ritcher. 1952. Biology of Deltochilum gibbosum (Fab.) with a description of the larva. The Coleopterists Bulletin 6: 53-57.
  7. Townes, H. and H. Howden. 1952. On the genotype of Geotrupes, and a discussion of the Fabrician 1798 usage of other names proposed by Latreille in 1796. Bulletin of Zoological Nomenclature 6: 207-209.
  8. Howden, H.F. 1954. A review of the genus Podolasia Harold (Coleoptera: Scarabaeidae). American Museum Novitates, No. 1661: 1-11.
  9. Olson, A.L., T.H. Hubbell and H.F. Howden. 1954. The burrowing beetles of the genus Mycotrupes (Coleoptera: Scarabaeidae: Geotrupinae). Miscellaneous Publication No. 84. University of Michigan Museum of Zoology: 1-50.
  10. Howden, H.F. 1954. Notes on Australian beetles in the tribe Bolboceratini formerly in the genus Bolboceras. Proceedings of the Linnean Society, New South Wales, 79: 142-144.
  11. Howden, H.F. 1955. The biology and taxonomy of the North American beetles of the subfamily Geotrupinae with revisions of the genera Bolbocerosoma, Eucanthus, Geotrupes, and Peltotrupes (Scarabaeidae). Proceedings of the United States National Museum 104: 151-319.
  12. Howden, H.F. 1955. Cases of interspecific "parasitism" in Scarabaeidae (Coleoptera). Journal of the Tennessee Academy of Science 31: 64-66.
  13. Howden, H.F. 1955. Description of a new Peruvian Athyreus with notes on the method of illustration. Entomologisches Arbeiten Museum Georg Frey, 6: 667-673.
  14. Howden, H.F. 1955. Some new species and records of North American Scarabaeidae (Coleoptera). Proceedings of the Entomological Society of Washington 57: 257-264.
  15. Howden, H.F., G.L. Cartwright and G. Halffter. 1956. Description de una nueva especie Mexicana de Onthophagus con anotaciones ecologicas sobre especies asociadas a nidos de animales y a cuevas. Acta Zoologica Mexicana 1(9): 1-16.
  16. Howden. H.F. 1957. Investigations on sterility and deformities of Onthophagus (Coleoptera, Scarabaeidae) induced by gamma radiation. Annals of the Entomological Society of America 50: 1-9.
  17. Howden. H.F. and P. Vaurie. 1957. Two new species of Trox from Florida (Scarabaeidae). American Museum Novitates 1818: 1-6.
  18. Howden, H.F. and S.I. Auerbach. 1953. Some effects of gamma radiation on Trogoderma sternale Jayne. Annals of the Entomological Society of America 51: 48-51.
  19. Howden, H.F. 1958. A seventh species of Podolasia Harold (Coleoptera: Scarabaeidae). The Canadian Entomologist 90: 291-292.
  20. Howden, H.F. 1958. The species of Acoma Casey having a three-segmented antennal club (Coleoptera: Scarabaeidae). The Canadian Entomologist 90: 337-401.
  21. Howden, H.F. 1959. A new species of Isonychus Mannerheim from southern Arizona (Coleoptera: Scarabaeidae). The Canadian Entomologist 91: 59-61.
  22. Howden, H.F. 1959. A new species of Anoplodera Mulsant in the mutabilis-aspera group (Coleoptera: Cerambycidae). The Canadian Entomologist 91: 157-160.
  23. Howden, H.F. 1959. Descriptions of two new species of Cyrtinus LeConte; with a key to the New World Cyrtinini (Coleoptera: Cerambycidae). The Canadian Entomologist 91: 372-375.
  24. Howden, H.F. 1960. Two new species of Cyrtinus LeConte (Coleoptera: Cerambycidae). The Canadian Entomologist 92: 173-177.
  25. Howden, H.F. 1960. A new species of Phyllophaga from the Big Bend region of Texas and Coahuila, with notes on other Scarabaeidae of the area. The Canadian Entomologist 92: 457-464.
  26. Becker, E.C., W.J., and H.F. Howden. 1960. Coleoptera in the Canadian National Collection. The Coleopterists Bulletin 14: 19-20.
  27. Crossley, D.A., Jr., and H.F. Howden. 1961. Insect-vegetation relationships in an area contaminated by radioactive wastes. Ecology 42: 302-317.
  28. Howden, H.F. and D.A. Crossley, Jr. 1961. Insect species on vegetation of the White Oak Lake bed, Oak Ridge, Tennessee. Oak Ridge, Tennessee. Oak Ridge National Laboratory 3094-UC-48. Biology and Medicine. 38 pp.
  29. Howden, H.F. 1961. New species and a new genus of Melolonthinae from the southeastern United States (Coleoptera: Scarabaeidae). The Canadian Entomologist 93: 807-812.
  30. Howden, H.F. 1961. A revision of the New World species of Thalycra Erichson, with a description of a new genus and notes on generic synonymy (Coleoptera: Nitidulidae). Memoirs of the Entomological Society of Canada No. 25: 61 pp.
  31. Howden, H.F. 1962. New species and synonymy in Acoma (Coleoptera: Scarabaeidae). The Canadian Entomologist 94: 1147-1155.
  32. Howden, H.F. 1962. The habits of some North American Scarabaeidae (Coleoptera). Proceedings of the Royal Entomology Society of London, Series C, Journals of Meetings 27: 17.
  33. Howden, H.F. and O.L. Cartwright. 1963. Scarab beetles of the genus Onthophagus Latreille north of Mexico (Coleoptera: Scarabaeidae). Proceedings of the United States National Museum 114: 1-135.
  34. Howden, H.F. and A. Martínez. 1963. The new tribe Athyreini and its included genera (Coleoptera: Scarabaeidae, Geotrupinae). The Canadian Entomologist 95: 345-352.
  35. Howden. H.F. 1963. Speculations on some beetles, barriers, and climates during the Pleistocene and pre-Pleistocene periods in some non-glaciated portions of North America. Systematic Zoology 12: 178-201.
  36. Howden, H. F. 1964. The Mercure trial: a side line of entomology. The Canadian Entomologist 96: 121.
  37. Howden, H.F. 1964. The Geotrupinae of North and Central America. Memoirs of the Entomological Society of Canada 39: 1-91.
  38. Howden, H.F. 1965. Notes on the Geotrupes of Taiwan (Formosa) with a description of a new species. Pacific Insects 7: 502-504.
  39. Howden. H.F. 1965. Lectotype designations in Cotinis Burm. for species described by Bates in the Biologica Centrali-Americana (Coleoptera: Scarabaeidae). The Canadian Entomologist 97: 670-671.
  40. Howden, H.F. 1965. A second New World species of Sisyphus Latreille (Coleoptera: Scarabaeidae). The Canadian Entomologist 97: 842-844.
  41. Howden, H.F. 1965. (Participant in) A symposium on factors affecting the distribution of plants and animals in northern Canada. Arctic Circular 16: 31-32.
  42. Howden, H. F. 1966. Entomological ramblings in Mexico. The Coleopterists Bulletin 20: 19-26.
  43. Howden, H.F. and S. Endrödi. 1966. Five new species of Cyclocephala Latreille from North and Central America (Coleoptera: Scarabaeidae). The Canadian Entomologist 98: 295-302.
  44. Howden, H.F. 1966. Notes on Canthonini of the "Biologia Centrali-Americana" and descriptions of new species (Coleoptera: Scarabaeidae). The Canadian Entomologist 98: 725-741.
  45. Howden, H.F. 1966. Some possible effects of the Pleistocene on the distributions of North American Scarabaeidae (Coleoptera). The Canadian Entomologist 98: 1177-1190.
  46. Howden. H.F. 1967. Mexican Geotrupini: a new species of Geotrupes and description of the larva of Ceratotrupes (Coleoptera: Scarabaeidae). The Canadian Entomologist 99: 1003-1007.
  47. Howden, H.F. 1968. A review of the Trichiinae of North and Central America (Coleoptera: Scarabaeidae). Memoirs of the Entomological Society of Canada 54: 1-77.
  48. Howden, H.F., H.E. Evans, and E.O. Wilson. 1968. A suggested revision of nomenclatural procedure in animal taxonomy. Systematic Zoology 13: 188-191.
  49. Howden, H.F. 1968. Generic relationships of Thyce, Plectrodes, Dinacoma and Hypotrichia with a description of a new genus and species from eastern Texas (Coleoptera: Scarabaeidae: Melolonthini). The Canadian Entomologist 100: 542-548.
  50. Howden. H.F. 1968. Canadian Ochodaeus, with a description of a new species. The Canadian Entomologist 100: 1118-1120.
  51. Howden, H.F. 1969. Effects of the Pleistocene on North American insects. Annual Review of Entomology 14: 39-56.
  52. Howden, H.F. 1969. Species of Coenonycha Horn from Mainland Baja, California, Mexico (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 23: 53-56.
  53. Howden, H.F. 1968(1969). Effects of man on the Ontario insect fauna. Proceedings of the Entomological Society of Ontario 99: 51-53.
  54. Howden, H.F. 1970. Jamaican Scarabaeidae: notes and descriptions (Coleoptera). The Canadian Entomologist 102: 1-15.
  55. Howden, H. F. 1970. First South Dakota records for Amblychila cylindriformis. Cicindela 2(3): 8.
  56. Howden, H.F. 1970. A new name for Onthophagus monticolus Howden and Cartwright. Proceedings of the Entomological Society of Washington 72: 54.
  57. Howden, H.F. 1970. Jamaican Cyrtinus, with descriptions of two new species (Coleoptera: Cerambycidae). The Canadian Entomologist 102: 1312-1316.
  58. Howden, H.F. 1970. The Coleoptera; in Fauna of Sable Island and its Zoogeographic Affinities. National Museum of Natural Sciences, Publications in Zoology, No. 4, pp. 1-30.
  59. Howden, H.F. 1970. The genus Paragnorimus, with descriptions of two new species (Coleoptera: Scarabaeidae). The Canadian Entomologist 102: 1385-1389.
  60. Howden, H.F. 1971. A second species of Peltotrichius (Coleoptera: Scarabaeidae: Trichiinae). The Canadian Entomologist 103: 104-106.
  61. Howden, H.F. 1971. Key to the New World Cremastocheilini, with notes and description of a new genus (Coleoptera: Scarabaeidae). Proceedings of the Entomological Society of Washington 73: 224-230.
  62. Howden, H.F. 1971. Five unusual genera of New World Scarabaeidae (Coleoptera). The Canadian Entomologist 103: 1463-1471.
  63. Howden, H.F. and A.R. Hardy. 1971. Generic placement and adult behaviour of the genus Leptohoplia Saylor (Coleoptera: Scarabaeidae). Proceedings of the Entomological Society of Washington 73: 337-341.
  64. Howden, H.F. 1972. New species of Dialithus Parry and a new synonym of Pantodinus Burmeister (Coleoptera: Scarabaeidae: Trichiinae). The Canadian Entomologist 104: 647-654.
  65. Howden, H.F. and S.B. Peck. 1972. Collecting in El Salvador, with particular reference to Scarabaeoidea and Silphoidea (Coleoptera). The Coleopterists Bulletin 26: 63-72.
  66. Howden. H.F. 1972. Systematics and zoogeography: science or politics? Systematic Zoology 21: 129-131.
  67. Kaiser, G.W., L.P. Lefkovitch, and H.F. Howden. 1972. Faunal provinces in Canada as exemplified by mammals and birds: a mathematical consideration. Canadian Journal of Zoology 50: 1087-1104.
  68. Howden, H.F. and L.E.C. Ling. 1973. Scanning Electron Microscopy: Low-magnification pictures of uncoated zoological specimens. Science 179: 386-388.
  69. Howden. H.F. 1973. Scarabatermes amazonensis, a new genus, new species of termitophilous Scarabaeidae (Coleoptera). The Canadian Entomologist 105: 29-33.
  70. Howden, H.F. 1973. A revision of the New World genus Cryptocanthon Balthasar (Coleoptera, Scarabaeidae). Canadian Journal of Zoology 51: 39-48.
  71. Gordon, R.D. and H.F. Howden. 1973. Five new species of Mexican Aphodius (Coleoptera: Scarabaeidae) associated with Thomomys umbrinus (Geomyidae). Annals of the Entomological Society of America 66: 436-443.
  72. Howden, H.F. 1973. Four new species of Onthophagus from Mexico and the United States. Proceedings of the Entomological Society of Washington 75: 329-337.
  73. Howden, H.F. 1973. Bolbothyreus, a new genus for Stenaspidius ruficollis (Coleoptera: Scarabaeidae: Geotrupinae). The Canadian Entomologist 105: 1567-1571.
  74. Howden, H.F. 1973. New species of Cyrtinus LeConte from Mexico and Venezuela (Coleoptera: Cerambycidae). The Canadian Entomologist 105: 595-597.
  75. Howden, H.F. 1974. A revision of the Australian genus Stenaspidius Westwood (Coleoptera, Scarabaeidae, Geotrupinae). Records of the South Australian Museum, 17: 11-21.
  76. Howden, H.F. and J.M. Campbell. 1974. Observations on some Scarabaeoidea in the Columbian Sierra Nevada de Santa Marta. The Coleopterists Bulletin 28: 109-114.
  77. Howden, H.F. 1974. Additional records and descriptions of North and Central American Geotrupinae (Coleoptera, Scarabaeidae). Canadian Journal of Zoology 52: 567-573.
  78. Howden, H.F. 1974. Problems in interpreting dispersal of terrestrial organisms as related to continental drift. Biotropica, 6: 1-6.
  79. Howden, H.F. 1974. Problems of dispersal of terrestrial organisms in relation to continental drift (reprinted from Biotropica). The American Association of Petroleum Geologists Bulletin 58: 2497-2501.
  80. Howden, H.F. and J.F. Lawrence. 1974. The New World Aesalinae, with notes on the North American lucanid subfamilies (Coleoptera, Lucanidae). Canadian Journal of Zoology 52: 1505-1510.
  81. Howden, H.F. and L.E.C. Ling. 1974. Low-magnification study of uncoated specimens, pp. 149-158. In: Principles and techniques of scanning electron microscopy (M.A. Hayat, ed.). Van Nostrand Reinhold Co., New York and Toronto.
  82. Howden, H.F., and V.G. Nealis. 1975. Effects of clearing in a tropical rain forest on the composition of the coprophagous scarab beetle fauna (Coleoptera). Biotropica, 7: 77-83.
  83. Howden, H.F. 1975. Late Quaternary history of some Maritime insects, pp. 61-65. In: Environmental Change in the Maritimes (J.F. Ogden III and M.J. Harvey, eds.). Nova Scotian Institute of Science, Halifax.
  84. Parry, R.H. and H.F. Howden. 1975. A new species of Coleopterus from Florida (Coleoptera, Nitidulidae). The Coleopterists Bulletin 29: 275-279.
  85. Howden, H.F. 1976. New species in the genera Bdelyropsis, Cryptocanthon and Drepanocerus (Coleoptera: Scarabaeidae). Proceedings of the Entomological Society of Washington 78: 95-103.
  86. Howden, H.F. 1976. A sixth species of Stenaspidius Westwood with a note on adult behaviour (Coleoptera: Scarabaeidae: Geotrupinae). Journal of the Australian Entomological Society 14: 439-441.
  87. Helava, J. and H.F. Howden. 1977. The genus Onthophilus Leach in Australia, with description of a new species (Coleoptera: Histeridae). Journal of the Australian Entomological Society 16: 83-86.
  88. Howden. H.F. 1977. Beetles, beach drift and island biogeography. Biotropica 9(1): 53-57.
  89. Howden, H.F. and J.B. Cooper. 1977. The generic classification of Bolboceratini of the Australian Region, with descriptions of four new genera (Scarabaeidae, Geotrupinae). Australian Journal of Zoology, Supplementary Series 50: 1-50.
  90. Howden, H.F. 1978. Descriptions of some West Indian Scarabaeidae primarily in the Natural History Museum, Basel. Entomologica Basiliensia, 3: 377-393.
  91. Howden, H.F. and A. Martinez. 1978. A review of the New World genus Athyreus Macleay (Scarabaeidae, Geotrupinae, Athyreini). Contributions of the American Entomological Institute 15(4): 1-70.
  92. Howden, H.F. and V.G. Nealis. 1978. Observations on height of perching in some tropical dung beetles (Scarabaeidae). Biotropica, 10: 43-46.
  93. Howden, H.F. 1979. A revision of the Australian genus Blackburnium Bouc. (Coleoptera: Scarabaeidae: Geotrupinae). Australian Journal of Zoology, Supplementary Series 72: 1-88.
  94. Howden, H.F. 1979. Scarabaeoidea, pp. 371-372. In: Canada and its insect fauna (H.V. Danks, ed.). Memoirs of the Entomological Society of Canada 108.
  95. Howden, H.F. 1980. Gillaspytes janzeni, a new genus, new species of North American Dynastinae (Coleoptera: Scarabaeidae), The Canadian Entomologist 112: 89-92.
  96. Howden, H.F. 1980. Athyreus Macleay, 1819, and Glyptus Brullé, 1835 (Insecta: Coleoptera): Proposed conservation Z.N. (S.) 1583. Bulletin of Zoological Nomenclature, 37: 191-192.
  97. Howden, H.F. 1980. Key to the Geotrupini of Mexico and Central America, with the description of a new species (Scarabaeidae, Geotrupinae). Canadian Journal of Zoology 58: 1959-1963.
  98. Howden. H.F. 1982. Zoogeography of some Australian Coleoptera as exemplified by the Scarabaeoidea, chapter 35, 29 pp. In: Ecological Biogeography of Australia (A. Keast, ed.). W. Junk publisher.
  99. Howden, H.F. 1981. Random and infrequent dispersals, pp. 188-194. In: Vicariance Biogeography (G. Nelson and D.E. Rosen, eds.). Columbia University Press, N.Y.
  100. Howden, H.F. and O.P. Young. 1981. Panamanian Scarabaeinae taxonomy, distribution, and habits. Contributions of the American Entomological Institute 18: 1-204.
  101. Howden, A.T. and H.F. Howden. 1981. The larva and adult biology of Rhinosimus viridiaeneus. The Canadian Entomologist 113: 1055-1060.
  102. Miller, S., R. D. Gordon, and H.F. Howden. 1981. Reevaluation of Pleistocene scarab beetles from Rancho La Brea, California (Coleoptera: Scarabaeidae). Proceedings of the Entomological Society of Washington 83(4): 625-630.
  103. Paulian, R. and H.F. Howden. 1982. Un nouveau genre de Ceratocanthides des Antilles (Col. Scarabaeoidea). Bulletin de la Société entomologique de France 87: 78-85.
  104. Howden, H.F. 1982. Larval and adult characters of Frickius Germain, its relationship to the Geotrupini, and a phylogeny of some major taxa in the Scarabaeoidea (Insecta: Coleoptera). Canadian Journal of Zoology 60: 2713-2724.
  105. Howden, H.F. 1983. Megasoma elephas (Cornizuelo, Rhinoceros Beetle), pp. 735-736. In: Costa Rican Natural History. (D.H. Janzen, ed.). University of Chicago Press, Chicago and London.
  106. Howden, H.F. 1983. Dichotomius carolinus colonicus (Rueda Caca, Dung Beetle), pp. 713-714. In: Costa Rican Natural History. (D.H. Janzen, ed.) University of Chicago Press, Chicago and London.
  107. Howden, H.F. and B. Gill. 1984. Two new species of Neoathyreus Howden and Martinez from Costa Rica with notes on other Athyreini from Mexico and Central America (Coleoptera: Geotrupinae). The Canadian Entomologist 116: 1637-1641.
  108. Howden, H.F. 1984. "Family: Scarabaeidae Subfamily: Geotrupinae." In: A Catalog of the Coleoptera of America, United States Department of Agriculture, Agriculture Handbook Number 529-34a: 1-17.
  109. Chalumeau, F. and H.F. Howden. 1984. Le Genre Euparia (Coleoptera: Scarabaeoidea: Aphodiinae) Saint-Fargeau et Serville, 1828. Bulletin de la Société Entomologique de Suisse 57: 85-94.
  110. Peck, S.B. and H.F. Howden. 1984. Response of a dung beetle guild to different sizes of dung bait in a Panamanian rainforest. Biotropica 16: 235-283.
  111. Howden. H.F. 1984. The Australian Blackburnium harslettae-triceratops complex (Scarabaeidae, Geotrupinae). The Coleopterists Bulletin 38(2): 190-191
  112. Gill, B.D. and H.F. Howden. 1985. A review of the North American genus Aphonus Leconte (Coleoptera: Scarabaeidae: Dynastinae). The Coleopterists Bulletin 39(2): 119-129.
  113. Howden, H.F. 1985. A revision of the South American genus Parathyreus Howden and Martinez (Coleoptera: Scarabaeidae: Geotrupinae). The Coleopterists Bulletin 39(2): 161-173.
  114. Howden, H.F. 1985. A revision of the South American species in the genus Neoathyreus Howden and Martinez (Coleoptera, Scarabaeidae, Geotrupinae). Contributions of the American Entomological Institute 21(4): 1-95.
  115. Peck, S.B. and H.F. Howden. 1985. Biogeography of scavenging scarab beetles in the Florida Keys: Post-Pleistocene landbridge islands. Canadian Journal of Zoology 63: 2730-2737.
  116. Howden, H.F. 1985. Species richness of geotrupine beetles (Scarabaeidae: Geotrupinae). In: Soil and litter invertebrates of some Australian-Mediterranean-type ecosystems (J.D. Majer and P. Greenslade, eds). WAIT Biology Bulletin 12: 14-15.
  117. Howden, H.F. 1985. Expansion and contraction cycles, endemism and area: the taxon cycle brought full circle, pp. 473-487. In: Taxonomy, Phylogeny and Zoogeography of Beetles and Ants (George E. Ball, ed.). W. Junk Publishers, Dordrecht. 514 pp.
  118. Howden, H.F. 1985. A revision of the Australian beetle genera Bolboleaus Howden and Cooper, Blackbolbus Howden and Cooper, and Bolborhachium Boucomont (Scarabaeidae: Geotrupinae). Australian Journal of Zoology, Supplementary Series 111: 1-179.
  119. Helava, J.V.T., H.F. Howden, and A.J. Ritchie. 1985. A review of the New World genera of the myrmecophilous and termitophilous subfamily Hetaeriinae (Coleoptera: Histeridae). Sociobiology, 10(2): 127-386.
  120. Howden, H.F. and C.H. Scholtz. 1986. Changes in a Texas dung beetle community between 1975 and 1985: (Scarabaeidae, Scarabaeinae). The Coleopterists Bulletin 40: 313-316.
  121. Scholtz, C.H. and H.F. Howden. 1987. A revision of the African Canthonina (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of the Entomological Society of Southern Africa 50: 75-119.
  122. Scholtz, C.H. and H.F. Howden. 1987. A revision of the Southern African genus Epirinus Reiche (Scarabaeidae: Scarabaeinae). Journal of the Entomological Society of Southern Africa 50: 121-154.
  123. Howden, H.F. and C.H. Scholtz. 1987. A revision of the African genus Odontoloma Boheman (Scarabaeidae: Scarabaeinae). Journal of the Entomological Society of Southern Africa 50: 155-192.
  124. Howden, H.F. and S.B. Peck. 1987. Adult habits, larval morphology and phylogenetic placement of Taurocerastes patagonicus Philippi (Scarabaeidae: Geotrupinae). Canadian Journal of Zoology 65: 329-332.
  125. Howden, H.F. and B. Gill. 1987. New species and new records of Panamanian and Costa Rican Scarabaeinae (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 41: 201-224.
  126. Howden, H.F. and B.D. Gill. 1988. Xenocanthus, a new genus of inquiline Scarabaeidae from southeastern Venezuela (Coleoptera). Canadian Journal of Zoology 66: 2071-2076.
  127. Howden, H.F. and B.D. Gill. 1988. A fourth genus of unusually modified Ceratocanthinae (Coleoptera: Scarabaeidae) from South America. Canadian Journal of Zoology 66: 2077-2079.
  128. Howden, H.F. and C.H. Scholtz. 1988. A second species of the African genus Peckolus Scholtz and Howden (Scarabaeidae: Scarabaeini). Journal of the Entomological Society of Southern Africa 51(1): 133-136.
  129. Howden, H.F. 1988. A new genus and four new species of New World Trichiinae (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 42(3): 241-250.
  130. Howden, H.F. 1989. The Geotrupinae of New Guinea (Coleoptera: Scarabaeidae). Invertebrate Taxonomy 3: 261-289.
  131. Howden, H.F. and B.C. Ratcliffe. 1990. An unusual new species of Trigonopeltastes Burmeister from Chiapas, Mexico (Coleoptera: Scarabaeidae: Trichiinae). The Canadian Entomologist 122(1/2): 1-4.
  132. Howden, H.F., Howden, A.T., and R.I. Storey. 1991. Nocturnal perching of Scarabaeine dung beetles (Coleoptera, Scarabaeidae) in an Australian tropical rain forest. Biotropica 23(1): 51-57.
  133. Morón, M.A. and H.F. Howden. 1992. A second species of Plesiosternus Morón with notes on other Heterosternina (Coleoptera: Scarabaeidae: Rutelinae). The Coleopterists Bulletin 46(1): 15-19.
  134. Morón, M.A. and H.F. Howden. 1992. A new species of Plusiotis Burmeister (Coleoptera: Scarabaeidae, Rutelinae) from Guatemala, C.A. The Canadian Entomologist 124(2): 205-210.
  135. Howden, H.F. 1992. A revision of the Australian beetle genera Eucanthus Westwood, Bolbobaineus Howden and Cooper, Australobolbus Howden and Cooper, and Gilletinus Boucomont (Scarabaeidae: Geotrupinae). Invertebrate Taxonomy 6: 605-717.
  136. Howden, H.F. 1992. First Canadian record of the subfamily Ceratocanthinae (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 46(3): 273.
  137. Howden, H.F., and R.I. Storey. 1992. Phylogeny of the Rhyparini and the new tribe Stereomerini, with descriptions of new genera and species (Coleoptera; Scarabaeidae; Aphodiinae). Canadian Journal of Zoology 70: 1810-1823.
  138. Howden, H.F. and B.D. Gill. 1993. A second species of Termitoderus from Angola and Namibia (Scarabaeidae: Aphodiinae: Termitoderini). Sociobiology 21(1): 203-208.
  139. Stebnicka, Z., and Howden, H.F. 1994. A revision of the Australian genus Podotenus A. Schmidt (Coleoptera: Scarabaeoidea: Aphodiini). Invertebrate Taxonomy 8: 17-62.
  140. Howden, H.F. 1993. New Bolboceratini from Western Australia (Scarabaeidae, Geotrupinae). Journal of the Australian Entomological Society 32: 379-386.
  141. Howden, H.F. and B.D. Gill. 1993. Mesoamerican Onthophagus Latreille in the dicranius and mirabilis species groups (Coleoptera: Scarabaeidae). The Canadian Entomologist 125:1091-1114.
  142. Howden, H.F. 1994 A new Geotrupes Latreille (Coleoptera: Scarabaeidae) from the Sierra de las Minas Range of Guatemala. The Coleopterists Bulletin 48(4): 345-348.
  143. Howden, H.F. 1994. A new Mexican Apeltastes Howden and notes on Guatemalan Paragnorimus Becker (Scarabaeidae, Trichiinae). The Coleopterists Bulletin 48(4): 394-353.
  144. Howden, H.F. and B.D. Gill. 1995. Trachycrusus, A new genus of Ceratocanthinae (Coleoptera, Scarabaeidae) with two new species from Peru. The Canadian Entomologist 127: 587-593.
  145. Howden, H.F. 1995. New Rhyparini from Fiji and the Philippines (Coleoptera: Scarabaeidae: Aphodiinae). The Coleopterists Bulletin 49(1): 23-27.
  146. Bordat, P. and H.F. Howden. 1995. Trois nouveaux genres, trois nouvelles espèces de Stereomerinae de Bornéo (Coleoptera, Aphodiidae). Bulletin de la Société entomologique de France 100(1): 11-20.
  147. Stebnicka, Z.T. and H.F. Howden. 1995. Revision of the Australian genera in the tribes Aphodiini, Aegialiini and Proctophanini (Coleoptera: Scarabaeidae: Aphodiinae). Invertebrate Taxonomy 9: 709-766.
  148. Howden, H.F. and A. Solis. 1995. Two new species of Geotrupinae (Coleoptera: Scarabaeidae) from Costa Rica. The Coleopterists Bulletin 49(3): 261-266.
  149. Cook, J., H.F. Howden, and S.B. Peck. 1995. The Galápagos Islands genus Neoryctes Arrow (Coleoptera: Scarabaeidae: Dynastinae). The Canadian Entomologist 127: 177-193.
  150. Stebnicka, Z.T. and H.F. Howden. 1996. Australian genera and species in the tribes Odontolochini, Psammodiini, Rhyparini, Stereomerini and part of the Eupariini (Coleoptera: Scarabaeoidea: Aphodiinae). Invertebrate Taxonomy 10: 97-107.
  151. Howden, H.F. 1996. West Indian Neoathyreus and their over-water dispersal (Coleoptera: Scarabaeidae: Geotrupinae). Journal of Natural History 30: 1503-1515.
  152. Storey, R.I. and H.F. Howden. 1996. Revision of Australoxenella Howden & Storey in Australia (Coleoptera: Scarabaeidae: Aphodiinae). Memoirs of the Queensland Museum 39(2): 365-380.
  153. Howden, H.F. 1997. Podolasiini Howden, new tribe, and a revision of the included genera, Podolasia Harold and Podostena Howden, new genus (Coleoptera: Scarabaeidae: Melolonthinae). The Coleopterists Bulletin 51(3): 223-255.
  154. Stebnicka, Z.T. and H.F. Howden. 1997. Revision of the Australian species of Ataenius Harold (Coleoptera: Scarabaeoidea: Aphodiinae: Eupariini). Invertebrate Taxonomy 11: 735-821.
  155. Howden, H.F. 1998. A new species of Pelidnota Macleay from Chiriqui Province, Panama (Scarabaeidae, Rutelinae). The Coleopterists Bulletin 52(2): 171-173.
  156. Howden, H.F. and L.J. Joly. 1998. South American Trigonopeltastes Burmeister (Coleoptera: Scarabaeidae), with description of a new species from Venezuela. The Coleopterists Bulletin 52(2): 194-200.
  157. Howden, H.F. 1998. More on Podolasia Harold: a new species and notes on another (Coleoptera: Scarabaeidae: Melolonthinae). The Coleopterists Bulletin 52(3): 275-277.
  158. Génier, F. and H.F. Howden. 1999. Two New Central American Onthophagus Latreille of the mirabilis species group (Coleoptera: Scarabaeidae: Scarabaeinae). The Coleopterists Bulletin 53(2): 130-144.
  159. Howden, H.F. 1999. New species of Central and South American Athyreini (Coleoptera: Scarabaeidae: Geotrupinae). The Coleopterists Bulletin 53(4): 339-354.
  160. Howden, H.F. and B.D. Gill. 2000. Tribes of New World Ceratocanthinae, with keys to genera and descriptions of new species (Coleoptera: Scarabaeidae). Sociobiology 35(2b): 281-329.
  161. Delgado, L. and H.F. Howden. 2000. Una especie nueva braquiptera de Onthophagus de Mexico (Coleoptera: Scarabaeidae: Scarabaeinae). Folia Entomologica Mexicana 109: 35-41.
  162. Howden, H.F. and R.I. Storey. 2000. New Stereomerini and Rhyparini from Australia, Borneo and Fiji (Coleoptera: Scarabaeidae: Aphodiinae). Memoirs of the Queensland Museum 46: 175-182.
  163. Gill, B.D. and H.F. Howden. 2000. Systematics and biology of the New World Ceratocanthinae (Coleoptera: Scarabaeidae). XXI- International Congress of Entomology, Brazil. Abstract Book I, Session 21 [3669].
  164. Morón, M.A. and H.F. Howden. 2001. New species in the genus Dilophochila Bates (Coleoptera: Melolonthidae: Rutelinae: Anomalini). The Coleopterists Bulletin 55: 51-64.
  165. Howden, H.F. 2001. A new tribe of Hybosorinae with a description of a new species of Callosides Howden (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 55: 199-204.
  166. Howden, H.F. and A.T. Howden. 2001. Change through time: a third survey of the Scarabaeinae (Coleoptera: Scarabaeidae) at Welder Wildlife Refuge. The Coleopterists Bulletin 55: 356-362.
  167. Howden, H.F. and A.T. Howden, 2001 (2000). Tetrops praeusta (L.) (Coleoptera: Cerambycidae), a potential pest? Insecta Mundi 14(4): 220.
  168. Howden, H.F. and B.D. Gill. 2001. A new species of Ivieolus Howden and Gill and a new type species designation for Aulisostes Howden and Gill. Sociobiology 38(3A): 531-538.
  169. Howden, H.F. and J. Cook. 2002. Paracryptocanthon, a new canthonine genus from Brazil (Coleoptera: Scarabaeidae: Scarabaeinae). The Coleopterists Bulletin 56: 585-588.
  170. Jameson, M.L. and H.F. Howden. 2002. Bolboceras Kirby, 1819 (July) (Insecta, Coleoptera): proposed precedence over Odonteus Samouelle, 1819 (June). Bulletin of Zoological Nomenclature 59: 246-248.
  171. Howden, H.F. 2002. The Athyreus tribuliformis Felsche complex with descriptions of three new species (Coleoptera: Scarabaeoidea: Geotrupidae: Athyreini). Entomotropica 17(1): 25-35.
  172. Howden, H.F. and B.D. Gill. 2003. The Mexican and Central American species of Astaenomoechus (Coleoptera, Scarabaeidae, Ceratocanthinae). Sociobiology 42: 201-242.
  173. Howden, H.F. 2003. Two new genera of New World Rhyparini (Coleoptera: Scarabaeidae: Aphodiinae). Sociobiology 42: 391-402.
  174. Howden, H.F. 2003. Subfamilia Geotrupinae, pp. 95-106. In: Atlas de los escarabajos de México. Coleoptera: Lamellicornia Volume 2 (M. A. Morón, ed.). Familias Scarabaeidae, Trogidae, Passalidae y Lucanidae. Argania Editio, Barcelona, Spain.
  175. Howden, H.F. 2003. Subfamilia Ceratocanthinae, pp. 114-121. In: Atlas de los escarabajos de México. Coleoptera: Lamellicornia Volume 2 (M. A. Morón, ed.). Familias Scarabaeidae, Trogidae, Passalidae y Lucanidae. Argania Editio, Barcelona, Spain.
  176. Jameson, M.L. and H.F. Howden. 2004. Comment on the proposed precedence of Bolboceras Kirby, 1819 (July) (Insecta, Coleoptera) over Odonteus Samouelle, 1819 (June). Bulletin of Zoological Nomenclature 61: 43-45.
  177. Howden, H.F. and F. Génier. 2004. Seven new species of Onthophagus Latreille from Mexico and the United States (Coleoptera: Scarabaeidae, Scarabaeinae). Fabreries 29: 53-76.
  178. Skelley, P.E. and H.F. Howden. 2005. A new species of Lomanoxia Martínez from Costa Rica (Coleoptera: Scarabaeidae: Aphodiinae). Insecta Mundi 17:185-190. [Dated 2003.]
  179. Howden, H.F. and B.D. Gill. 2005. An unusual new species of Germarostes Paulian from Peru (Coleoptera: Scarabaeidae: Ceratocanthinae). The Coleopterists Bulletin 59(2): 272-274.
  180. Howden, H.F. and V. Maly. 2005. Cryptoryctes tricornutus sp. nov. from the Northern Territory, Australia (Scarabaeidae, Dynastinae, Oryctini). The Canadian Entomologist 137: 274-277.
  181. Howden, H.F. 2005. Notes on Bolboceratini from northeastern Mexico with a description of a new Bolbocerosoma Schaeffer (Coleoptera: Scarabaeidae: Geotrupinae). Folia Entomologica Mexicana 44(2): 213-218.
  182. Howden, H.F. 2006. New species and a new subgenus of South American Neoathyreus Howden and Martínez (Coleoptera: Geotrupidae: Athyreini). Zootaxa 1137: 37-52.
  183. Howden, H.F. 2006. Two new species of Odontotrypes Fairmaire from northern Myanmar (Coleoptera: Geotrupidae). Acta Zoologica Cracoviensia 49B: 13-16.
  184. Howden, H.F. 2006. Case 3359. Germarostes Paulian, 1982 and Haroldostes Paulian, 1982 (Insecta, Coleoptera, Scarabaeidae): proposed conservation. Bulletin of Zoological Nomenclature 63(4): 239-242.
  185. Howden, H.F. and A. Smetana. 2007. Case 3377. Ataenius Harold, 1867 (Insecta, Coleoptera): proposed precedence over Aphodinus Motschulsky, 1862. Bulletin of Zoological Nomenclature 64(1): 39-42.
  186. Howden, H.F., A.T. Howden, and G. Holloway. 2007. Digging down under: Australian Bolboceratini, their habits and a list of species (Coleoptera: Scarabaeoidea: Geotrupidae). Zootaxa 1499: 47-59.
  187. Howden, H.F. 2007. In past years – I. Scarabs 19: 17-18.
  188. Howden, H.F. 2007. In past years – II – Ecuador. Scarabs 20: 23-25.
  189. Howden, H.F. 2007. In past years – III – The Australian “out-back”. Scarabs 21: 12-15.
  190. Howden, H.F. 2007. Strategus antaeus (Drury) notes on biology. Scarabs 22: 2-3.
  191. Howden, H.F. 2007. In past years – IV – Australia & Argentina. Scarabs 22: 9-14.
  192. Howden, H.F. 2008. In past years – V – with George Vogt. Scarabs 23: 9-12.
  193. Howden, H.F. 2008. In past years – VI – 1940’s – 50’s. Scarabs 24: 9-14.
  194. Howden, H.F. 2008. In past years – VII – late 1950’s – early 60’s. Scarabs 25: 5-19.
  195. Howden, H.F. 2008. In past years – VIII – Bernard Benesh. Scarabs 26: 14.
  196. Howden, H.F. 2008. In past years – IX 1964-69. Scarabs 27: 7-21.
  197. Howden, H.F. 2008. Association of Cetoniinae (Coleoptera: Scarabaeidae) with Thrincopyge LeConte (Coleoptera: Buprestidae) in North America. The Coleopterists Bulletin 62(2): 181-184.
  198. Howden, H.F. 2008. In past years – XI 1970-1974. Scarabs 28: 14-19.
  199. Howden, H.F. 2008. In past years – XII 1970-1974. Scarabs 29: 16-22.
  200. Howden, H.F. 2008. In past years – XIII 1970-1974. Scarabs 30: 15-21.
  201. Howden, H.F. 2008. In past years – XIV 1970-1974. Scarabs 31: 14-23.
  202. Howden, H.F. 2008. In past years – XV 1974-1975 sabbatical. Scarabs 32: 12-19.
  203. Howden, H.F. 2008. In past years – XVI – 1975. Scarabs 33: 14-21.
  204. Howden, H.F. 2008. In past years – XVII – 1975. Scarabs 34: 15-22.
  205. Howden, H.F. 2009. In past years – XVIII – 1976-1977. Scarabs 35: 15-21.
  206. Howden, H.F. 2009. In past years – XIX – 1977. Scarabs 36: 16-20.
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