Checklist of the Scarabaeoidea of southern South America
Andrew B. T. Smith
Canadian Museum of Nature, P.O. Box 3443, Station D
Ottawa, ON, K1P 6P4, CANADA
asmith@mus-nature.ca
Disclaimer: the nomenclature and classification used in this checklist are based on what has been published in the scientific literature. It should be noted that this includes numerous errors, omissions, misclassifications, and misidentifications that have not yet been rectified. Additionally, there are a number of new taxa I am aware of that have not yet been published. Corrections will be made to this checklist as taxonomic revisions and reviews are published. In the mean time, users of the information provided in this checklist need to be cautious!
This checklist is a work in progress. A very small percentage of the entries and refences have not yet been verified using the original literature.
Southern South America consists of the IV-XII Regions of Chile and the following provinces in Argentina: Neuquén, Río Negro, Chubut, Santa Cruz, and Tierra del Fuego.
Type genus: Lucanus Scopoli, 1763: 1.
Type genus: Aesalus Fabricius, 1801: 254.
Type genus: Ceratognathus Westwood, 1838: 260
Genus Hilophyllus Paulsen and Mondaca, 2006
Hilophyllus Paulsen and Mondaca, 2006: 6.
Type species: Ceratognathus penai Martínez, 1976 by original designation.
Hilophyllus argentinensis (Martínez, 1981)
Original combination: Ceratognathus argentinensis Martínez, 1981: 13.
Type locality: "Argentina, provincia de Neuquén, Lago Curruhe Grande."
Distribution: Argentina: Neuquén, Chile: VIII-X Región (Paulsen and Mondaca 2006).
Hilophyllus martinezi Paulsen and Mondaca, 2006
Original combination: Hilophyllus martinezi Paulsen and Mondaca, 2006: 16.
Type locality: "CHILE ÑUBLE / Los Lleuques."
Distribution: Chile: VIII Región del Biobío (Paulsen and Mondaca 2006).
Hilophyllus penai (Martínez, 1976)
Original combination: Ceratognathus penai Martínez, 1976: 101.
Type locality: "Chile, Arauco, Nahuelbutá (W), Caramavida."
Distribution: Chile: IX Región de la Araucania (Martínez 1976).
Subfamily LAMPRIMINAE MacLeay, 1819
Type genus: Lamprima Latreille, 1807: 132.
Genus Streptocerus Fairmaire, 1850
Streptocerus Fairmaire, 1850: 53.
Type species: Streptocerus speciosus Fairmaire 1850 by monotypy.
Streptocerus speciosus Fairmaire, 1850
Original combination: Streptocerus speciosus Fairmaire, 1850: 55.
Type locality: "Chili."
Distribution: Argentina: Neuquén, Río Negro; Chile: VI Región del O'Higgins, X Región de Los Lagos (Solier 1851, Philippi and Philippi 1864, Bruch 1911).
Synonym: Streptocerus dejeanii Solier, 1851: 44.
Type locality: "la provincia de Valdivia."
Synonym: Streptocerus eustictus Philippi and Philippi, 1864: 316.
Type locality: "subandinis prov. Colchaguae" (Chile).
Synonym: Streptocerus regius Kriesche, 1922: 118.
Type locality: "Chile, Parral."
Type genus: Lucanus Scopoli, 1763: 1.
Genus Apterodorcus Arrow, 1943
Apterodorcus Arrow, 1943: 139.
Type species: Lucanus bacchus Hope, 1845 by monotypy.
Apterodorcus bacchus (Hope in Westwood, 1845)
Original combination: Lucanus bacchus Hope, 1845: 26 (Westwood, 1845).
Type locality: "Chiloe."
Distribution: Argentina: Chubut, Neuquén, Río Negro, Chile: VII Regin de Maule to X Región de Los Lagos (Philippi 1859, Bruch 1911, Mondaca and Paulsen 2008).
Synonym: Dorcus bipunctatus Philippi, 1859: 656.
Type locality: "la provincia de Valdivia."
Apterodorcus tristis (Deyrolle in Parry, 1870)
Original combination: Sclerostomus tristis Deyrolle, 1870: 95 (Parry 1870).
Type locality: "Chili."
Distribution: Chile: VI Región de O'Higgins, RM Regin de Metropolitana (Mondaca and Paulsen 2008).
Genus Pycnosiphorus Solier, 1851
Pycnosiphorus Solier, 1851: 56.
Type species: Dorcus mandibularis Solier, 1851 by monotypy.
Synonym: Epipedus Solier, 1851: 49.
Type species: Lucanus caelatus Blanchard, 1842 by subsequent designation (Maes 1992).
Synonym: Erichius Maes, 1992: 52.
Type species: Dorcus femoralis Guérin-Méneville, 1839 by original designation.
Synonym: Calvertius Maes, 1992: 53.
Type species: Sclerostomus calverti Nagel, 1932 by original designation.
Synonym: Neoepipedus Özdikmen and Turgut, 2006: 145.
Type species: Lucanus caelatus Blanchard, 1842 by original designation (replacement name for Epipedus Solier, 1851).
Pycnosiphorus arriguttii Weinreich, 1958
Original combination: Pycnosiphorus arriguttii Weinreich, 1958: 172.
Type locality: "Las Cubras, Chile."
Distribution: Chile.
Pycnosiphorus brevicollis Benesh, 1956
Original combination: Pycnosiphorus brevicollis Benesh, 1956: 62.
Type locality: "Constitucion, Costa de Maule, Chile."
Distribution: Chile: VII Región del Maule, X Región de Los Lagos (Benesh 1956).
Pycnosiphorus calverti (Nagel, 1932)
Original combination: Sclerostomus calverti Nagel, 1932: 116.
Type locality: "Cunco, Chile."
Distribution: Chile: IX Región de La Araucania (Nagel 1932).
Synonym: Pycnosiphorus ariasi Weinreich, 1958: 169.
Type locality: "Chile, Cherquenco."
Pycnosiphorus caelatus (Blanchard, 1842)
Original combination: Lucanus caelatus Blanchard, 1842: plate 12.
Type locality: "Chili."
Distribution: Argentina: Neuquén; Chile (Blanchard 1847, Bruch 1911).
Synonym: Lucanus variolosus Hope, 1845: 25 (Westwood 1845).
Type locality: "Chiloe."
Pycnosiphorus costatus (Benesh, 1956)
Original combination: Sclerostomus costatus Benesh, 1956: 65.
Type locality: No data.
Distribution: Chile: VIII Región del Biobío, IX Región de La Araucania (Benesh 1956).
Pycnosiphorus fairmairii (Parry, 1864)
Original combination: Sclerostomus fairmairii Parry, 1864: 61.
Type locality: "Chili."
Distribution: Argentina: Neuquén; Chile (Parry 1864, Bruch 1911).
Pycnosiphorus fasciatus (Germain, 1855)
Original combination: Dorcus fasciatus Germain, 1855: 397.
Type locality: "Concepcion."
Distribution: Chile: VII Región del Biobío (Germain 1855).
Pycnosiphorus femoralis (Guérin-Méneville, 1839)
Original combination: Dorcus femoralis Guérin-Méneville, 1839: 303.
Type locality: "côtes du détroit de Magellan."
Distribution: Argentina: Neuquén, Río Negro, Chubut, Santa Cruz; Chile: XII Región de Magallanes (Curtis 1845, Bruch 1911).
Synonym: Dorcus darwinii Hope, 1841: 302.
Type locality: "Chili."
Synonym: Dorcus rufifemoralis Curtis, 1844: 200.
Type locality: "Port Famine."
Synonym: Lucanus rubripes Hope, 1845: 26 (Westwood 1845).
Type locality: "Magellan."
Synonym: Dorcus leiocephalus Solier, 1851: 53.
Type locality: "en la República [Chile]."
Synonym: Dorcus rufipes Solier, 1851: 50.
Type locality: "varias provincias de la República" [Chile].
Synonym: Dorcus spinolae Solier, 1851: 52.
Type locality: "en la República [Chile]."
Synonym: Dorcus modestus Philippi, 1859: 657.
Type locality: "la provincia de Valdivia."
Pycnosiphorus franzae Weinreich, 1958
Original combination: Pycnosiphorus franzae Weinreich, 1958: 173.
Type locality: "Contulmo, Mittelchile, Prov. Concepcion."
Distribution: Chile: VIII Región del Biobío.
Pycnosiphorus germaini Numhauser, 1985
Original combination: Pycnosiphorus germaini Numhauser, 1985: 71.
Type locality: "Cordillera de Malleco y Cordillera de Curacautín" (not specified).
Distribution: Chile: IX Región de La Araucania (Numhauser 1985).
Pycnosiphorus lessonii lessonii (Buquet, 1842)
Original combination: Dorcus lessonii Buquet, 1842: 283.
Type locality: "Chili."
Distribution: Chile (Buquet 1842).
Pycnosiphorus lessonii marginipennis Deyrolle, 1870
Original combination: Pycnosiphorus marginipennis Deyrolle, 1870: 95 (Parry 1870).
Type locality: "Chili."
Distribution: Chile (Parry 1870).
Synonym: Pycnosiphorus pulcher Landin, 1955: 4.
Type locality: "Estero Pichi Pilluco, E of Puerto Montt."
Pycnosiphorus magnificus Benesh, 1956
Original combination: Pycnosiphorus magnificus Benesh, 1956: 60.
Type locality: "Tregualemu, Chile."
Distribution: Chile: Tregualemu (Benesh 1956).
Pycnosiphorus mandibularis (Solier, 1851)
Original combination: Dorcus mandibularis Solier, 1851: 56.
Type locality: "en la Repúblic [Chile]."
Distribution: Chile: VII Región del Maule, XI Región Aisen (Numhauser 1985).
Pycnosiphorus pallidocinctus (Fairmaire and Germain, 1861)
Original combination: Dorcus pallidocinctus Fairmaire and Germain, 1861: 2.
Type locality: "Chilensia."
Distribution: Chile (Fairmaire and Germain 1861).
Pycnosiphorus philippi (Westwood, 1864)
Original combination: Sclerostomus philippi Westwood, 1864: 61 (Parry 1864).
Type locality: "Chili."
Distribution: Argentina: Neuquén (Roig-Juñent et al. 2005); Chile.
Synonym: Sclerostomus elongatus Deyrolle, 1870: 96 (Parry 1870).
Type locality: "Chili."
Pycnosiphorus rubrovittatus (Blanchard, 1842)
Original combination: Lucanus rubrovittatus Blanchard, 1842: plate 12.
Type locality: "Chili."
Distribution: Chile (Blanchard 1847).
Synonym: Dorcus rouleti Solier, 1851: 53.
Type locality: "Chile."
Pycnosiphorus virgatus (Nagel, 1928)
Original combination: Sclerognathus virgatus Nagel, 1928: 290.
Type locality: "Tolhuaca."
Distribution: Chile: IX Región de La Araucania (Nagel 1928).
Pycnosiphorus vittatus (Eschscholtz, 1822)
Original combination: Lucanus vittatus Eschscholtz, 1822: 9.
Type locality: "Chili, Conception."
Distribution: Argentina: Neuquén; Chile: Región del Biobío (Eschscholtz 1822, Bruch 1911).
Pycnosiphorus westwoodi Numhauser, 1985
Original combination: Pycnosiphorus westwoodi Numhauser, 1985: 71.
Type locality: "desde Curicó (35º) a Valdivia 39º46')" (not specified).
Distribution: Chile: VI Región del Maule, X Región de Los Lagos (Numhauser 1985).
Genus Sclerostomus Burmeister, 1847
Sclerostomus Burmeister, 1847: 423 (replacement name for Sclerognathus [Westwood], 1845).
Type species: Sclerognathus costatus [Westwood], 1845 by monotypy for Sclerognathus [Westwood], 1845.
Synonym: Sclerognathus [Westwood], 1845: 26.
Type species: Sclerognathus costatus [Westwood], 1845 by monotypy.
Remark: junior homonym of Sclerognathus Valenciennes, 1844 (Pisces).
Sclerostomus varasi Nagel, 1932
Original combination: Sclerostomus varasi Nagel, 1932: 115.
Type locality: "Termae de Chillan (Las termas de Chillan)."
Distribution: Chile: VII Región del Maule, VIII Región del Biobío (Paulsen 2005).
Genus Sclerostomus (Chileistomus) Weinreich, 1960
Sclerostomus (Chileistomus) Weinreich, 1960: 74.
Type species: Lucanus cucullatus Blanchard, 1842 by original designation.
Sclerostomus (Chileistomus) cucullatus (Blanchard, 1842)
Original combination: Lucanus cucullatus Blanchard, 1842: plate 12.
Type locality: "Chili."
Distribution: Argentina: Neuquén; Chile: Región VI-X (Paulsen 2005).
Genus Sclerostomus (Sclerostomulus) Weinreich, 1960
Sclerostomus (Sclerostomulus) Weinreich, 1960: 88.
Type species: Dorcus tuberculatus Solier, 1851 by original designation.
Sclerostomus (Sclerostomulus) nitidus Benesh, 1955
Original combination: Sclerostomus nitidus Benesh, 1955: 99.
Type locality: No data.
Distribution: Chile: VI Región del Bernardo O'Higgins (Paulsen 2005).
Sclerostomus (Sclerostomulus) tuberculatus (Solier, 1851)
Original combination: Dorcus tuberculatus Solier, 1851: 54.
Type locality: "la Araucania."
Distribution: Chile: Región VII-X (Paulsen 2005).
Genus Bomansodus Chalumeau and Brochier, 1995
Synonym: Bomansodus Chalumeau and Brochier, 1995: 20.
Type species: Chiasognathus impubis Parry, 1870 by original designation.
Bomansodus impubis (Parry, 1870)
Original combination: Chiasognathus impubis Parry, 1870: 68.
Type locality: "Chili... Mendoza, on the eastern side of the Cordillera."
Distribution: Argentina: Mendoza; Chile: IX Región de la Araucania, X Región de Los Lagos (Parry 1870, Chalumeau and Brochier 2007).
Genus Chiasognathus Stephens, 1831
Chiasognathus Stephens, 1831: 213.
Type species: Chiasognathus grantii Stephens, 1831 by original designation.
Synonym: Tetropthalma Lesson, 1833: 5.
Type species: Tetropthalma chiloensis Lesson, 1833 by monotypy.
Synonym: Carmenia Molino-Olmedo, 2001: 85.
Type species: Chiasognathus latreillei Solier, 1851 by monotypy.
Remark: Junior homonym of Carmenia Roewer, 1915.
Synonym: Ramirezia Molino-Olmedo, 2002: 124.
Type species: Chiasognathus jousellinii Reiche, 1850 by original designation.
Remark: Junior homonym of Ramirezia Zamponi, 1979.
Synonym: Carmeniella Molino-Olmedo, 2003: 29.
Type species: Chiasognathus latreillei Solier, 1851 by original designation.
Remark: Replacement name for Carmenia Molino-Olmedo, 2001.
Synonym: Ramireziella Molino-Olmedo, 2003: 29.
Type species: Chiasognathus jousellinii Reiche, 1850 by original designation.
Remark: Replacement name for Ramirezia Molino-Olmedo, 2002.
Chiasognathus beneshi Lacroix, 1979
Original combination: Chiasognathus beneshi Lacroix, 1979: 250.
Type locality: "Chili, Province d'Aysen: Rio Manihuales."
Distribution: Chile: VIII Región del Biobío, XI Región Aisen (Lacroix 1979, Chalumeau and Brochier 2007).
Chiasognathus grantii Stephens, 1831
Original combination: Chiasognathus grantii Stephens, 1831: 214.
Type locality: "Island of Chiloe."
Distribution: Argentina: Neuquén, Río Negro; Chile: VIII-XI Región (Bruch 1911, Chalumeau and Brochier 2007).
Synonym: Tetropthalma chiloensis Lesson, 1833: 6.
Type locality: "l'île de Chiloë."
Synonym: Chiasognathus affinis Philippi, 1859: 658.
Type locality: "cerca del Corral."
Synonym: Chiasognathus pygmaeus Dallas, 1933: 74.
Type locality: not verified
Chiasognathus jousellinii Reiche, 1850
Original combination: Chiasognathus jousellinii Reiche, 1850: 249 (Reiche 1850a).
Type locality: "versant oriental des Andes du Chili" (Reiche 1850b).
Distribution: Argentina, Chile: VIII-XI Región (Reiche 1850b, Chalumeau and Brochier 2007).
Synonym: Chiasognathus mniszechii Thomson, 1862: 406.
Type locality: "Chili."
Synonym: Chiasognathus schoenemanni Kriesche, 1919: 94.
Type locality: "Chile, Bäder von Longavi, Parral."
Chiasognathus latreillei Solier, 1851
Original combination: Chiasognathus latreillei Solier, 1851:42.
Type locality: "Chile."
Distribution: Argentina: Neuquén, Río Negro, Chubut; Chile: IX-XI Región (Chalumeau and Brochier 2007).
Synonym: Chiasognathus imberbis Philippi, 1859: 657.
Type locality: "cerca de la ciudad de Valdivia."
Synonym: Chiasognathus reichei Thomson, 1862: 407.
Type locality: "Chili."
Type genus: Geotrupes Latreille, 1797: 6.
Type genus: Geotrupes Latreille, 1797: 6.
Type genus: Taurocerastes Philippi, 1866: 115.
Genus Frickius Germain, 1897
Frickius Germain, 1897: 294.
Type species: Frickius variolosus Germain, 1897 by subsequent designation (Zunino 1984).
Frickius costulatus Germain, 1897
Original combination: Frickius costulatus Germain, 1897: 299.
Type locality: "las Trancas, punto situado tres leguas antes de llegar a las Termas de Chillan."
Distribution: Chile: VIII Región del Biobío (Germain 1897).
Frickius variolosus Germain, 1897
Original combination: Frickius variolosus Germain, 1897: 296.
Type locality: "provenia de Valdivia... volcan Villarrica."
Distribution: Argentina Neuquén, Chubut, Santa Cruz; Chile: VIII Región del Biobío, X Región de Los Lagos, XI Región Aisen (Bruch 1911, Gutiérrez 1947, 1949).
Genus Taurocerastes Philippi, 1866
Taurocerastes Philippi, 1866: 115.
Type species: Taurocerastes patagonicus Philippi, 1866 by monotypy.
Taurocerastes patagonicus Philippi, 1866
Original combination: Taurocerastes patagonicus Philippi, 1866: 116.
Type locality: "Patagonia haud procul a colonia chilensi in freto Magellanico sita."
Distribution: Argentina: Chubut, Santa Cruz; Chile: XI Región Aisen, XII Región de Magallanes; Falkland Islands (Gutiérrez 1947, Schweiger 1959).
Type genus: Bolboceras Kirby, 1819: 459.
Type genus: Bolboceras Kirby, 1819: 459.
Genus Bolborhinum Boucomont, 1911
Bolborhinum Boucomont, 1911: 339.
Type species: Bolboceras tubericeps Fairmaire, 1856 by original designation.
Bolborhinum binasutum (Fairmaire and Germain, 1861)
Original combination: Bolboceras binasutum Fairmaire and Germain, 1861: 2
Type locality: "Chilensia."
Distribution: Chile: VIII Región del Bíobío, IX Región de la Araucania (Gutiérrez 1949).
Bolborhinum geotrupoides (Laporte, 1840)
Original combination: Bolboceras geotrupoides Laporte, 1840: 104.
Type locality: "Chili."
Distribution: Chile (Laporte 1840).
Synonym: Bolboceras distinguendum Fairmaire and Germain, 1861: 2 (emended from Bolboceras distinguendus).
Type locality: "Chilensia."
Bolborhinum laesicolle (Fairmaire, 1856)
Original combination: Bolboceras laesicolle Fairmaire, 1856: 483 (emended from Bolboceras laesicollis).
Type locality: "Chilensia."
Distribution: Argentina: Neuquén (Martínez 1976); Chile: IX Región de la Araucania, X Región de Los Lagos (Gutiérrez 1949).
Synonym: Bolboceras excavatum Philippi, 1859: 660 (emended from Bolboceras excavatus).
Type locality: "cerca de Corral."
Synonym: Bolboceras mundum Redtenbacher, 1868: 60.
Type locality: "Chili."
Bolborhinum nasutum (Fairmaire and Germain, 1861)
Original combination: Bolboceras nasutum Fairmaire and Germain, 1861: 2.
Type locality: "Chilensia."
Distribution: Chile: VI Región del Bernardo O'Higgins (Gutiérrez 1949).
Synonym: Bolboceras andicola Philippi, 1873: 312.
Type locality: "Andibus prov. Santiago."
Bolborhinum shajovskoyi (Martínez, 1952)
Original combination: Bolboceras shajovskoyi Martínez, 1952: 318.
Type locality: "Argentina, territorio nacional de Neuquén, San Martín de los Andes."
Distribution: Argentina: Neuquén (Martínez 1952).
Bolborhinum tricorne (Solier, 1851)
Original combination: Bolboceras tricorne Solier, 1851: 67 (emended from Bolboceras tricornis).
Type locality: "la provincia de Santiago."
Distribution: Chile: V-VII Región (Gutiérrez 1949).
Bolborhinum tubericeps (Fairmaire, 1856)
Original combination: Bolboceras tubericeps Fairmaire, 1856: 483.
Type locality: "Chilensia."
Distribution: Chile: VII Región del Maule, VIII Región del Biobío (Gutiérrez 1949).
Synonym: Bolboceras bicorne Philippi, 1859: 659 (emended from Bolboceras bicornis).
Type locality: "en nuestro potrero de San-Juan" (Valdivia).
Synonym: Bolboceras tetraodon Redtenbacher, 1868: 59.
Type locality: "Chili."
Genus Zefevazia Martínez, 1954
Zefevazia Martínez, 1954: 193 (Martínez 1954b).
Type species: Zefevaziarosascostai Martínez, 1954 by original designation.
Zefevazia cantisanii (Martínez, 1976)
Original combination: Bolborhinum (Zefevazia) cantisanii Martínez, 1976: 535.
Type locality: "la provincia de Buenos Aires, Partido de Puán, Estación Filipe Solá."
Distribution: Argentina: Catamarca, Córdoba, Buenos Aires, San Rafael, La Rioja, Mendoza, Río Negro (Martínez 1976).
Type genus: Glaresis Erichson, 1848: 925.
Genus Glaresis Erichson, 1848
Glaresis Erichson, 1848: 925.
Type species: Glaresis rufa Erichson, 1848 by monotypy.
Glaresis pardoalcaidei Martínez, Pereira, and Vulcano, 1961
Original combination: Glaresis pardoalcaidei Martínez, Pereira, and Vulcano 1961: 69.
Type locality: "cuidad de San Miguel del Tucumán, Puente del río Sali."
Distribution: Brasil; Bolivia; Paraguay; Argentina: Salta, Catamarca, Tucumán, Chaco, Santiago del Estero, Córdoba, La Rioja (Scholtz 1990).
Glaresis fritzi Martínez, Pereira, and Vulcano, 1961
Original combination: Glaresis fritzi Martínez, Pereira, and Vulcano 1961: 77.
Type locality: "provincia de Río Negro, Lamarque."
Distribution: Argentina: Catamarca, Río Negro (Scholtz 1990).
Type genus: Trox Fabricius, 1775: 31.
Remark: Keys to genera in Scholtz (1990).
Genus Trox Fabricius, 1775
Trox Fabricius, 1775: 31.
Type species: Scarabaeus sabulosus Linnaeus, 1758 by subsequent designation (Latreille 1810).
Remark: Key to species in Scholtz (1990).
Synonym: Pseudotrox Robinson, 1948: 1.
Type species: Trox laticollis LeConte, 1854 by original designation.
Trox scaber (Linnaeus, 1767)
Original combination: Silpha scabra Linnaeus, 1767: 573.
Type locality: "Europa."
Distribution: Palaearctic; Nearctic; Australia; Argentina: Corrientes, Buenos Aires; Chile: V-VI Región (Vaurie 1962, Scholtz 1990).
Synonym: Trox barbosus von Laicharting, 1781: 31.
Type locality: "Tyroler."
Synonym: Trox arenarius Fabricius, 1787: 18.
Type locality: "Halae Saxonum."
Synonym: Trox arenosus Gmelin, 1790: 1586.
Type locality: "Halae Saxonum."
Synonym: Trox hispidus von Paykull, 1798: 81.
Type locality: "Suecica."
Synonym: Trox trisulcatus Curtis, 1844: 197.
Type locality: "Valparaiso."
Synonym: Trox niponensis Lewis, 1895: 387.
Type locality: "St. Paul's Island in the Indian Ocean and... Hakodate [Japan]."
Genus Omorgus (Omorgus) Erichson, 1847
Omorgus Erichson, 1847: 111 (Erichson 1847a).
Type species: Trox suberosus Fabricius, 1775 by subsequent designation.
Remark: Key to species in Scholtz (1990).
Synonym: Chesas Burmeister, 1876: 264.
Type species: Trox pastillarius Blanchard, 1847 by monotypy.
Synonym: Lagopelas Burmeister, 1876: 265.
Type species: Trox ciliatus Blanchard, 1847 by monotypy.
Synonym: Megalotrox Preudhomme De Borre, 1886: 59.
Type species: Trox gigas Harold, 1872 by subsequent designation.
Omorgus (Omorgus) candezei (Harold, 1872)
Original combination: Trox candezei Harold, 1872: 113.
Type locality: "Pampas von Südamerika."
Distribution: Argentina: San Luis, Santiago del Estero (Scholtz 1990).
Omorgus (Omorgus) ciliatus (Blanchard, 1847)
Original combination: Trox ciliatus Blanchard, 1847: 190.
Type locality: "village de Patagones en Patagonie."
Distribution: Bolivia; Argentina: Salta, Catamarca, Santiago del Estero, Mendoza, Buenos Aires, Neuquén, Río Negro (Scholtz 1990).
Omorgus (Omorgus) nocheles Scholtz, 1990
Original combination: Omorgus (Omorgus) nocheles Scholtz, 1990: 1412.
Type locality: "Argentina: El Bolsón, Rio Negro."
Distribution: Argentina: Río Negro (Scholtz 1990).
Omorgus (Omorgus) pastillarius (Blanchard, 1847)
Original combination: Trox pastillarius Blanchard, 1847: 187.
Type locality: "en Patagonie, seulement dans les dunes près de la mer, en dehors de la baie de San-Blas."
Distribution: Argentina: Salta, Tucumán, Catamarca, Santiago del Estero, La Rioja, Entre Ríos, Córdoba, San Luís, Mendoza, Buenos Aires, Neuquén, Chubut (Blanchard 1847, Scholtz 1990).
Remark: The country records for Bolivia and Chile in Vaurie (1962) are dubious. Further study is needed before these countries can be added to the distributional range of this species.
Omorgus (Omorgus) persuberosus (Vaurie, 1962)
Original combination: Trox persuberosus Vaurie, 1962: 145.
Type locality: "Ypiranga, São Paulo, Brazil."
Distribution: Perú; Bolivia; Brazil; Paraguay; Uruguay; Argentina: Chaco, Tucumán, Santiago del Estero, Mendoza, Buenos Aires, Neuquén (Vaurie 1962, Scholtz 1990).
Omorgus (Omorgus) suberosus (Fabricius, 1775)
Original combination: Trox suberosus Fabricius, 1775: 31.
Type locality: "Brasilia."
Distribution: North America; Central America; West Indes; Venezuela; Guiana, French Guiana, Colombia; Ecuador; Perú; Bolivia; Brazil; Paraguay; Uruguay; Argentina: Jujuy, Salta, Tucumán, Chaco, Catamarca, Santiago del Estero, La Rioja, Córdoba, Entre Ríos, San Luis, Buenos Aires, Río Negro, Chubut, Santa Cruz; Chile: Atacama; (Vaurie 1962, Scholtz 1990).
Synonym: Trox crenatus Olivier, 1789: 7 (Olivier 1789b).
Type locality: "Cayenne."
Synonym: Trox gibbus Olivier, 1789: 13 (Olivier 1789b).
Type locality: "Saint-Domingue."
Synonym: Trox ovatus Palisot de Beauvois, 1818: 175.
Type locality: "Saint-Domingue."
Synonym: Trox denticulatus Palisot de Beauvois, 1818: 176.
Type locality: "États-Unis d'Amérique; Pensylvanie."
Synonym: Trox alternatus Say, 1835: 179.
Type locality: "Pennsylvania... Arkansaw... New Orleans."
Synonym: Trox nobilis Wollaston, 1867: 93.
Type locality: "Porto da Luz, in Fogo" (Cape Verde Archipelago).
Synonym: Trox tricolor Blackburn, 1904: 295.
Type locality: "New South Wales."
Synonym: Trox manilensis Schultze, 1916: 295.
Type locality: "Luzon, Manila."
Synonym: Trox torressalai Báguena, 1959: 212.
Type locality: "España... Alicante, Denia."
Synonym: Omorgus triestinae Pittino, 1987: 378.
Type locality: "Brésil Minas-Geraes."
Genus Polynoncus Burmeister, 1876
Polynoncus Burmeister, 1876: 264.
Type species: Trox pedestris Harold, 1872 by subsequent designation (Scholtz 1982).
Remark: Key to species in Scholtz (1990).
Polynoncus aeger (Guérin-Méneville, 1830)
Original combination: Trox aeger Guérin-Méneville, 1830: plate 22.
Type locality: "Pérou."
Distribution: Brazil; Uruguay; Argentina: Jujuy, Catamarca, Corrientes, Entre Ríos, Buenos Aires, Neuquén, Río Negro; Chile: VI-VIII Región (Vaurie 1962, Scholtz 1990).
Synonym: Trox leprosus Blanchard, 1847: 188.
Type locality: "Maldonado et à Montevideo."
Polynoncus brevicollis (Eschscholtz, 1822)
Original combination: Trox brevicollis Eschscholtz, 1822: 12.
Type locality: "Chili, Conception."
Distribution: Perú; Argentina: Buenos Aires, Neuquén, Chubut, Río Negro; Chile: IV-X Región (Gutiérrez 1949, Scholtz 1990).
Remark: Locality records from Colombia, Paraguay, and Perú (Scholtz 1990) are dubious and should be removed from the known distribution of this species.
Synonym: Trox lachrymosus Curtis, 1844: 197.
Type locality: "Valparaiso."
Polynoncus bullatus (Curtis, 1844)
Original combination: Trox bullatus Curtis, 1844: 197.
Type locality: "Valparaiso."
Distribution: Argentina: Neuquén; Chile: III-VIII Región, X Región de Los Lagos (Scholtz 1990).
Polynoncus chilensis (Harold, 1872)
Original combination: Trox chilensis Harold, 1872: 137.
Type locality: "Chili" and "San Jago."
Distribution: Argentina: Río Negro; Chile: VII Región del Maule, IX-XI Región (Scholtz 1990).
Polynoncus diffluens (Vaurie, 1962)
Original combination: Trox diffluens Vaurie, 1962: 138.
Type locality: "Leyda, Santiago, Chile."
Distribution: Chile: VI Región del Bernardo O'Higgins (Scholtz 1990).
Polynoncus gemmifer (Blanchard, 1847)
Original combination: Trox gemmifer Blanchard, 1847: 187 (emended from Trox gemmiferus).
Type locality: "Patagonie" (Blanchard 1847).
Distribution: Uruguay; Argentina: San Luis, Córdoba, La Pampa, La Rioja, Catamarca, Buenos Aires, Mendoza, Río Negro, Salta, Santiago del Estero, Tucumán (Scholtz 1990).
Synonym: Trox argentinus Harold, 1872: 143.
Type locality: "Cordova im argentinischen Freistaat..., Buenos Aires."
Polynoncus gibberosus Scholtz, 1990
Original combination: Polynoncus gibberosus Scholtz 1990: 1428.
Type locality: "Chile: Concepcion."
Distribution: Argentina: Neuquén; Chile: VIII Región del Biobío, IX Región de La Araucania (Scholtz 1990).
Polynoncus guttifer (Harold, 1868)
Original combination: Trox guttifer Harold, 1868: 86 (Harold 1868c) (replacement name for Trox gemmiferus Blanchard, 1847: 188, a junior homonym of Trox gemmiferus Blanchard, 1847: 187).
Type locality: "Patagonie, dans les endroits élevés, secs et sablonneux, près le village de Patagones" (Blanchard 1847).
Distribution: Paraguay; Argentina: Mendoza, Buenos Aires, Neuquén, Río Negro, Chubut, Santa Cruz (Scholtz 1990).
Synonym: Trox gemmiferus Blanchard, 1847: 188 (a junior homonym of Trox gemmiferus Blanchard, 1847: 187).
Type locality: "Patagonie, dans les endroits élevés, secs et sablonneux, près le village de Patagones."
Polynoncus haafi (Vaurie, 1962)
Original combination: Trox haafi Vaurie, 1962: 122.
Type locality: "Golfe St. Georges, Coli-Huapi [Comodoro Rivadavia, Argentina]."
Distribution: Argentina: Neuquén, Río Negro, Chubut, Santa Cruz (Scholtz 1990).
Polynoncus hemisphaericus (Burmeister, 1876)
Original combination: Trox hemisphaericus Burmeister, 1876: 253.
Type locality: "Rio Sa Cruz im südlichen Patagonien."
Distribution: Argentina: Buenos Aires, Neuquén, Chubut, Río Negro, Santa Cruz; Chile: XII Región de Magallanes (Vaurie 1962, Scholtz 1990).
Synonym: Trox globulatus Fairmaire, 1884: 490.
Type locality: "Punta-Arena."
Polynoncus longitarsis (Harold, 1872)
Original combination: Trox longitarsis Harold, 1872: 136.
Type locality: "Chili."
Distribution: Argentina: Neuquén, Río Negro, Chubut; Chile: VIII-X Región (Scholtz 1990).
Polynoncus mirabilis Pittino, 1987
Original combination: Polynoncus mirabilis Pittino, 1987: 391.
Type locality: "Santiago Peron."
Distribution: Argentina: Río Negro, Chubut; Chile: VI Región del Bernardo O'Higgins, VIII-XI Región (Scholtz 1990).
Polynoncus neuquen (Vaurie, 1962)
Original combination: Trox neuquen Vaurie, 1962: 123.
Type locality: "Neuquen [Argentina]."
Distribution: Argentina: Neuquén (Scholtz 1990).
Remark: Dubious records from Coquimbo, Chile and Salta, Argentina were listed by Vaurie (1962) and Scholtz 1990 respectively.
Polynoncus patagonicus (Blanchard, 1847)
Original combination: Trox patagonicus Blanchard, 1847: 186.
Type locality: "en Patagonie... e'était à l'entrée du Rio Negro sur les dunes."
Distribution: Brazil; Argentina: Jujuy, Tucumán, Buenos Aires, Río Negro, Chubut, Santa Cruz (Vaurie 1962, Scholtz 1990).
Polynoncus pedestris (Harold, 1872)
Original combination: Trox pedestris Harold, 1872: 128.
Type locality: "Bei den Salinen von Andres-Paz in Patagonien...; Cordova im argentinischen Freistaat...; Chilecito."
Distribution: Argentina: Tucumán, Catamarca, San Luis, Santiago del Estero (Scholtz 1990).
Polynoncus pilularius (Germar, 1824)
Original combination: Trox pilularius Germar, 1824: 113.
Type locality: "Buenos Ayres."
Distribution: Perú; Bolivia; Brazil; Paraguay; Uruguay; Argentina: Jujuy, Salta, Chaco, Tucumán, Catamarca, Santiago del Estero, Entre Ríos, Buenos Aires, Neuquén (Scholtz 1990).
Type genus: Hybosorus MacLeay, 1819: 120.
Type genus: Ceratocanthus White, 1842: 93.
Type genus: Ceratocanthus White, 1842: 93.
Genus Germarostes (Germarostes) Paulian, 1982
Germarostes (Germarostes) Paulian, 1982: 13.
Type species: Melolontha aphodioides Illiger, 1800 by original designation.
Germarostes (Germarostes) posticus (Germar, 1843)
Original combination: Acanthocerus posticus Germar, 1843: 144.
Type locality: "Chili."
Distribution: Argentina, Chile: IV-VII Región (Gutiérrez 1946, 1949).
Synonym: Acanthocerus muricatus Curtis, 1844: 197.
Type locality: "Valparaiso."
Synonym: Cloeotus mochae Gutiérrez, 1946: 14.
Type locality: "Isla de La Mocha."
Genus Martinezostes Paulian, 1982
Martinezostes Paulian, 1982: 63.
Type species: Acanthocerus asper Philippi, 1859 by original designation.
Martinezostes asper (Philippi, 1859)
Original combination: Acanthocerus asper Philippi, 1859: 660.
Type locality: "la provincia de Valdivia."
Distribution: Chile: X Región de Los Lagos (Gutiérrez 1949).
Martinezostes fortecostatus (Gutiérrez, 1949)
Original combination: Cloeotus fortecostatus Gutiérrez, 1949: 9.
Type locality: "Provincia de Valdivia, Valdivia."
Distribution: Chile: X Región de Los Lagos (Gutiérrez, 1949).
Martinezostes ruizi (Gutiérrez, 1946)
Original combination: Cloeotus ruizi Gutiérrez, 1946: 13.
Type locality: "proveniente de Curicó. Zona maritima."
Distribution: Chile: VII Región del Maule (Gutiérrez 1946).
Type genus: Scarabaeus Linnaeus, 1758: 345.
Type genus: Aphodius Illiger, 1798: 15 (Kugelann and Illiger 1798).
Type genus: Aegialia Latreille, 1807: 96.
Remark: Keys to genera in Stebnicka et al. (2004).
Genus Aegialia (Aegialia) Latreille, 1807
Aegialia Latreille, 1807: 96.
Type species: Aphodius globosus Illiger, 1798 by monotypy.
Aegialia argentina Martínez, Pereira, and Vulcano, 1970
Original combination: Aegialia (Aegialia) argentina Martínez, Pereira, and Vulcano, 1970: 336.
Type locality: "Río Negro, San Antonio Oeste, Playa Las Grutas."
Distribution: Argentina: Catamarca, La Rioja, Río Negro (Martínez et al. 1970).
Genus Amerisaprus Stebnicka and Skelley, 2004
Amerisaprus Stebnicka and Skelley, 2004: 79 (in Stebnicka et al. 2004).
Type species: Amerisaprus valdivia Stebnicka and Skelley, 2004 by original designation.
Amerisaprus valdivia Stebnicka and Skelley, 2004
Original combination: Amerisaprus valdivia Stebnicka and Skelley, 2004: 79 (in Stebnicka et al. 2004).
Type locality: "Chile, Valdivia Prov., 34 km WNW La Union."
Distribution: Chile: X Región de Los Lagos (Stebnicka et al. 2004).
Genus Argeremazus Stebnicka and Dellacasa, 2004
Argeremazus Stebnicka and Dellacasa, 2004: 75 (in Stebnicka et al. 2004).
Type species: Argeremazus neuquen Stebnicka and Dellacasa, 2004 by original designation.
Argeremazus neuquen Stebnicka and Dellacasa, 2004
Original combination: Argeremazus neuquen Stebnicka and Dellacasa, 2004: 76 (in Stebnicka et al. 2004).
Type locality: "Argentina, Neuquen, Covunco."
Distribution: Argentina: Neuquén (Stebnicka et al. 2004).
Type genus: Proctophanes Harold, 1861: 111.
Genus Australaphodius Balthasar, 1942
Aphodius (Australaphodius) Balthasar, 1942: 203.
Type species: Aphodius melbournicus Balthasar, 1942 by monotypy.
Synonym: Phaeaphodius Reitter, 1892: 67.
Type species: Aphodius solskyi Harold, 1871 by original designation.
Australaphodius frenchi (Blackburn, 1892)
Original combination: Aphodius frenchi Blackburn, 1892: 35.
Type locality: "S. Australia and Victoria."
Distribution: Chile: V Región de Valparaiso, VIII-X Región; U.S.A.; South Africa; Australia; New Zealand (Dellacasa et al. 2002).
Synonym: Aphodius ambiguus Boheman, 1858: 279.
Type locality: not verified.
Synonym: Aphodius brevitarsis Péringuey, 1901: 388.
Type locality: "Cape Colony (Cape Town)" South Africa.
Synonym: Aphodius tarsalis Schmidt, 1907: 201.
Type locality: "Cape Colony (Cape Town)" South Africa.
Synonym: Aphodius melbournicus Balthasar, 1942: 203.
Type locality: "Süd-Australien."
Synonym: Aphodius catulus Balthasar, 1946: 54.
Type locality: "Africa austr., Caffraria, sine indicatione loci."
Type genus: Aphodius Illiger, 1798: 15 (Kugelann and Illiger 1798).
Genus Acanthaphodius Schmidt, 1909
Acanthaphodius Schmidt, 1909: 67 (Schmidt 1909c).
Type species: Acanthaphodius bruchi Schmidt, 1909 by monotypy.
Acanthaphodius bruchi Schmidt, 1909
Original combination: Acanthaphodius bruchi Schmidt, 1909: 67 (Schmidt 1909c).
Type locality: "Rio Negro."
Distribution: Argentina: Neuquén, Río Negro; Chile: IX Región de la Araucania, X Región de Los Lagos (Dellacasa and Stebnicka 2001).
Genus Aphodius (Calamosternus) Motschulsky, 1859
Calamosternus Motschulsky, 1859: 156.
Type species: Scarabaeus granarius Linnaeus, 1767 by monotypy.
Synonym: Megalisus Mulsant and Rey, 1870: 209 (Mulsant and Rey 1870a).
Type species: Aphodius stercorarius Mulsant and Rey, 1870 by subsequent designation (Dellacasa 1988).
Synonym: Pseudesymus d'Orbigny, 1896: 218.
Type species: Aphodius lucidus Klug, 1845 by monotypy.
Aphodius (Calamosternus) granarius (Linnaeus, 1767)
Original combination: Scarabaeus granarius Linnaeus, 1767: 547.
Type locality: "Europa."
Distribution: Nearctic; Palaearctic; Argentina: Buenos Aires, Chubut; Chile: V Región de Valparaíso, Región Metropolitana; Isla Juan Fernández (Schmidt 1931, Gutiérrez 1947).
Synonym: Scarabaeus quadrituberculatus Fabricius, 1798: 23.
Type locality: "America."
Synonym: Aphodius niger Creutzer, 1799: 20.
Type locality: not verified.
Synonym: Aphodius carbonarius Sturm, 1805: 128.
Type locality: not verified.
Synonym: Aphodius haemorrhous Stephens, 1830: 196.
Type locality: "Suffolk, and within the metropolitan district" (Britain).
Synonym: Aphodius lucens Stephens, 1830: 196.
Type locality: "within the metropolitan district, and in Norfolk" (Britain).
Synonym: Aphodius emarginatus Stephens, 1830: 198.
Type locality: "near London."
Synonym: Aphodius melanopus Stephens, 1830: 198.
Type locality: "throughout the metropolitan district; also in Suffolk and Norfolk" (Britain).
Synonym: Aphodius elongatus Ménétries, 1832: 182.
Type locality: "Bakou."
Synonym: Aphodius retusus Waltl, 1835: 67.
Type locality: not verified.
Synonym: Aphodius suturalis Faldermann, 1835: 259.
Type locality: "Persico-Armeniaca."
Synonym: Aphodius concolor Mulsant, 1842: 199.
Type locality: "France."
Synonym: Aphodius cribratus Mulsant, 1842: 199.
Type locality: "France."
Synonym: Aphodius parcepunctatus Mulsant, 1842: 199.
Type locality: "France."
Synonym: Aphodius rugosulus Mulsant, 1842: 199.
Type locality: "France."
Synonym: Aphodius aterrimus Melsheimer, 1846: 136.
Type locality: "Maryland" (United States).
Synonym: Aphodius adelaidae Hope, 1847: 284.
Type locality: "New Holland."
Synonym: Aphodius metallicus Haldeman, 1848: 105.
Type locality: "Maryland."
Synonym: Aphodius spretus Haldeman, 1848: 106.
Type locality: "Middle States" (U.S.A.).
Synonym: Aphodius hypocrita Mulsant and Rey 1870: 449 (Mulsant and Rey 1870b).
Type locality: "France."
Synonym: Aphodius thoracicus Dalla Torre, 1879: 107.
Type locality: "Oberösterreich" (Austria).
Synonym: Aphodius inutilis Horn, 1887: 50.
Type locality: "San Francisco, Cal., and in Oregon."
Synonym: Aphodius brunnescens Reitter, 1892: 51.
Type locality: "Araxes und Syrien."
Synonym: Aphodius ragusanus Reitter, 1906: 718.
Type locality: not verified.
Synonym: Aphodius nama Kolbe, 1908: 126.
Type locality: "Steinkopf."
Synonym: Aphodius signatus Schmidt, 1916: 99.
Type locality: "Oberösterreich" (Austria).
Synonym: Aphodius pirinensis Balthasar 1946: 62.
Type locality: "Bulgaria (Macedonia), Montes Pirin."
Synonym: Aphodius tilgranicus Nakane 1966: 232.
Type locality: not verified.
Genus Aphodius (Labarrus) Mulsant and Rey, 1870
Labarrus Mulsant and Rey, 1870: 516 (Mulsant and Rey 1870b).
Type species: Scarabaeus lividus Olivier, 1789 by monotypy.
Synonym: Pseudocalamosternus Balthasar, 1936: 1.
Type species: Pseudocalamosternus rigidus Balthasar, 1936 by monotypy.
Aphodius (Labarrus) pseudolividus Balthasar, 1941
Original combination: Aphodius pseudolividus Balthasar, 1941: 148.
Type locality: "America meridionale, di diverse località, specialmente dei dintorni di Buenos Aires, Fran Chaco, Paraguay."
Distribution: Nearctic; Palaearctic; Argentina: Tucumán, Misiones, Buenos Aires, Mendoza; Chile: V Región deValparaíso, XII Región de Magallanes, Isla de Pascua (Gutiérrez 1947, Campos and Peña 1973). (Catamarca, Cordoba, Atacama – UCDC)
Genus Orodaliscoides Schmidt, 1913
Orodaliscoides Schmidt, 1913: 145.
Type species: Aphodius rugosiceps Harold, 1859 by subsequent designation (in Dellacasa 1988).
Orodaliscoides reflexus (Schmidt, 1910)
Original combination: Aphodius reflexus Schmidt, 1910: 357.
Type locality: "Argentina."
Distribution: Argentina: Chubut (Bruch 1911).
Orodaliscoides rugosiceps (Harold, 1859)
Original combination: Aphodius rugosiceps Harold, 1859: 213.
Type locality: "Chili."
Distribution: Chile (Harold 1859).
Podotenus (Pseudopodotenus) Dellacasa, 1992
Aphodius (Pseudopodotenus) Dellacasa, 1992: 152.
Type species: Aphodius fulviventris Fairmaire and Germain, 1860 by original designation.
Podotenus (Pseudopodotenus) fulviventris (Fairmaire and Germain, 1860)
Original combination: Aphodius fulviventris Fairmaire and Germain, 1860: 268.
Type locality: "Chili."
Distribution: Chile: VIII Región del Biobío, IX Región de la Araucania (Dellacasa 1992).
Genus Symphodon Schmidt, 1913
Symphodon Schmidt, 1913: 128.
Type species: Aphodius anomalus Harold, 1874 by monotypy.
Symphodon anomalus (Harold, 1874)
Original combination: Aphodius anomalus Harold, 1874: 185.
Type locality: "Chili."
Distribution: Chile: VIII Región del Biobío (Dellacasa 1992).
Type genus: Euparia LePeletier and Serville, 1828: 357.
Genus Ataenius Harold, 1867
Ataenius Harold, 1867: 82.
Type species: Ataenius scutellaris Harold, 1867 by subsequent designation (Cartwright 1974).
Synonym: Aphodinus Motschulsky, 1862: 55.
Type species: Aphodius castanicolor Motschulsky, 1858 by subsequent designation (Dellacasa et al. 2001).
Synonym: Hexalus Mulsant and Rey, 1870: 200.
Type species: Hexalussimplicipes Mulsant and Rey, 1870 by monotypy.
Synonym: Pseudammoecius Schmidt, 1913: 160.
Type species: Aphodiusaustralasiae Boheman, 1858 by monotypy.
Synonym: Placopterus Chalumeau, 1983: 144.
Type species: Ataenius tuberculatus Schmidt 1911 by monotypy.
Ataenius chilensis (Solier, 1851)
Original combination: Aphodius chilensis Solier, 1851: 72.
Type locality: "Coquimbo y Santiago."
Distribution: Chile: V Región de Valparaíso, Región Metropolitana (Solier 1851).
Synonym: Ataenius modestus Schmidt, 1912: 199.
Type locality: "Chile."
Ataenius gracilis (Melsheimer, 1846)
Original combination: Oxyomus gracilis Melsheimer, 1846: 137.
Type locality: "Pennsylvania."
Distribution: Nearctic; Chile: V Región de Valparaíso, Región Metropolitana, X Región de Los Lagos (Gutiérrez 1947, Landin 1955).
Ataenius opatroides (Blanchard, 1847)
Original combination: Oxyomus opatroides Blanchard, 1847: 185.
Type locality: "dans les endroits sablonneux à Montevideo et à Maldonado, près de la mer."
Distribution: Uruguay; Argentina: Buenos Aires, Córdoba; Chile: Región Metropolitana, VII-X Región (Stebnicka 2004).
Ataenius picinus Harold, 1868
Original combination: Ataenius picinus Harold, 1868: 281 (Harold 1868a).
Type locality: "Chili."
Distribution: Oceania; Australia; New Zealand; Nearctic; Central America; West Indes; Bolivia; Brasil; Paraguay; Uruguay; Argentina: Corrietes, Entre Ríos, Formosa, Jujuy, Misiones, Santiago de Estero; Chile: VII Región del Maule, IX Región de la Araucania, X Región de Los Lagos (Stebnicka 2004).
Synonym: Ataenius duplopunctatus Lea, 1923: 6.
Type locality: "Western Australia: Parkerville."
Synonym: Ataenius salutator Fall, 1930: 99.
Type locality: "Pensacola, Florida."
Synonym: Ataenius queirosii Paulian, 1934: 219.
Type locality: "Vaté, Nouvelles-Hébrides."
Synonym: Ataenius darlingtoni Hinton, 1937: 179.
Type locality: "Porto Rico: Caratanga, Lagoon."
Synonym: Ataenius boucomontii Paulian, 1937: 41.
Type locality: "Australie: Sydney."
Synonym: Ataenius alegrus Balthasar, 1947: 50.
Type locality: "Brasilia, Porto Alegro."
Synonym: Saprosites rugosus Richards, 1959: 41.
Type locality: "Mt. Roskill, Auckland" (New Zealand).
Synonym: Ataenius paracognatus Balthasar, 1961: 123.
Type locality: "Bolivien, El Beni."
Ataenius platensis (Blanchard, 1847)
Oxyomus platensis Blanchard, 1847: 185.
Type locality: "Montevideo et Corrientes."
Distribution: Uruguay; Argentina: Buenos Aires, San Luis, Corrientes, Chaco, Santiago del Estero, Mendoza, Catamarca (Blanchard 1847, Bruch 1911).
Synonym: Ataenius integer Harold, 1868: 86 (Harold 1868b).
Type locality: "Brasilia."
Synonym: Ataenius anticus Fall, 1930: 105.
Type locality: "Hope, Arkansas."
Synonym: Ataenius granchaocensis Balthasar, 1939: 28.
Type locality: "Paraguay, Gran Chaco."
Synonym: Ataenius histrionicus Balthasar, 1947: 49.
Type locality: "Paraguay, prov. Alto Parana."
Synonym: Ataenius heyrovskyi Balthasar, 1960: 5.
Type locality: "Brasilien, Rio Madeira."
Synonym: Ataenius degallieri Chalumeau, 1990.
Type locality: not verified
Ataenius strigicaudus Bates, 1887
Original combination: Ataenius strigicaudus Bates, 1887: 96 (emended from Ataenius strigicauda).
Type locality: "Mexico, Cordova, Tuxtla, Jalapa; Guatemala, Paso Antonio; Nicaragua, Chontales; Panama. - South Brazil; Amazons; Antilles, St. Thomas."
Distribution: México; Central America; West Indes; Colombia; Ecuador; Perú; Bolivia; Brasil; Paraguay; Argentina: Misiones, La Rioja; Chile: VIII Región del Biobío (Stebnicka 2004).
Synonym: Ataenius aspericollis Petrovitz, 1973: 178.
Type locality: "Brasilien, Acre, Feijó... und Paraná, Foz de Iguassu."
Genus Bruchaphodius Martínez, 1952
Bruchaphodius Martínez, 1952: 99.
Type species: Euparia bruchi Schmidt, 1922 by original designation.
Bruchaphodius ovalipennis (Harold, 1871)
Original combination: Euparia ovalipennis Harold, 1871: 116.
Type locality: "Cordova Argentinae."
Distribution: Paraguay; Argentina: Santiago del Estero, Córdoba, Mendoza (Bruch 1911, Martínez 1952).
Bruchaphodius shannoni (Bruch, 1938)
Original combination: Euparia shannoni Bruch, 1938: 160.
Type locality: "Famabalasto, valle de Santa María (Catamarca)."
Distribution: Argentina: Catamarca (Bruch 1938).
Genus Oxyataenius Dellacasa and Stebnicka, 2001
Oxyataenius Dellacasa and Stebnicka, 2001: 30.
Type species: Oxyomus morosus Harold, 1869 by original designation.
Oxyataenius morosus (Harold, 1869)
Original combination: Oxyomus morosus Harold, 1869: 100 (Harold 1869b).
Type locality: "Chili."
Distribution: Chile (Dellacasa and Stebnicka 2001).
Genus Parataenius Balthasar, 1961
Parataenius Balthasar, 1961: 121.
Type species: Parataenius mirabilis Balthasar, 1961 by original designation.
Synonym: Brancoataenius Paulian, 1979: 66.
Type species: Brancoataenius lusitanicus Paulian, 1979 by original designation.
Parataenius simulator (Harold, 1868)
Original combination: Ataenius simulator Harold, 1868: 85 (Harold 1868b).
Type locality: "Mendoza."
Distribution: Nearctic; Palaearctic; Austalia; Bolivia; Brasil; Uruguay; Argentina: San Luis, La Rioja, Mendoza; Chile (Bruch 1911, Chalumeau 1992).
Synonym: Psammodius schwarzi Linell, 1896: 721.
Type locality: "Jacksonville, Florida."
Synonym: Parataenius granuliceps Petrovitz, 1971: 102.
Type locality: "N. S. W., Sidney."
Synonym: Ataenius lusitanicus Paulian, 1979: 66.
Type locality: "Portugal, Fos de Neiva, Minho."
Tribe PSAMMODIINI Schmidt, 1910
Type genus: Psammodius Fallén, 1807: 37.
Genus Leiopsammodius Rakovic, 1981
Leiopsammodius Rakovic, 1981: 16 (Rakovic 1981b).
Type species: Psammodius laevicollis Klug, 1845 by original designation.
Leiopsammodius indefensus (Schmidt, 1909)
Original combination: Psammobius indefensus Schmidt, 1909: 60 (Schmidt 1909b).
Type locality: "Valparaiso."
Distribution: Chile: V Región de Valparaíso (Cartwright 1955).
Synonym: Leiopsammodius chilensis Rakovic, 1990: 5.
Type locality: "Chile."
Leiopsammodius placidus (Schmidt, 1911)
Original combination: Psammobius placidus Schmidt, 1911: 38.
Type locality: "Argentina."
Distribution: Argentina (Schmidt 1911).
Genus Odontopsammodius Gordon and Pittino, 1992
Odontopsammodius Gordon and Pittino, 1992: 265.
Type species: Psammodius cruentus Harold, 1868 by original designation.
Odontopsammodius cruentus (Harold, 1868)
Original combination: Psammodius cruentus Harold, 1868: 282 (Harold 1868a).
Type locality: "chilensischen."
Distribution: Nearctic; Bolivia; Paraguay; Argentina: Buenos Aires, Córdoba, La Rioja, Mendoza, Misiones, Salta, San Juan, San Luis, Santiago del Estero, Tucumán; Chile: Región Metropolitana (Bruch 1911, Gutiérrez 1947, Cartwright 1955).
Remark: Cartwright (1955) doubted the Chilean record.
Synonym: Psammobius shermani Cartwright, 1946: 89.
Type locality: "Sullivans Island, South Carolina."
Genus Platytomus Mulsant, 1842
Platytomus Mulsant, 1842: 310.
Type species: Platytomus sabulosus Mulsant, 1842 by monotypy.
Platytomus micros (Bates, 1887)
Original combination: Psammodius micros Bates, 1887: 103.
Type locality: "Guatemala, Capetillo, Dueñas."
Distribution: Nearctic; Central America; Chile: Isla de Pascua (Campos and Peña 1973).
Synonym: Pleurophorus batesi Arrow, 1903: 514.
Type locality: "Mexico, Presidio..; Guatemala, Dueñas, Paraiso.."
Synonym: Diastictus californicus Petrovitz 1963: 645
Type locality: "Californien, S. Barbara."
Genus Pleurophorus Mulsant, 1842
Pleurophorus Mulsant, 1842: 312.
Type species: Scarabaeus caesus Panzer, 1796 by monotypy.
Pleurophorus caesus (Panzer, 1796)
Original combination: Scarabaeus caesus Panzer, 1796: 2.
Type locality: "sub lapidibus Viennae."
Distribution: Chile: Isla Juan Fernández (Schmidt 1931).
Synonym: Aphodius angustus Philippi and Philippi, 1864: 316.
Type locality: "sub lapidibus, in terra, nunquam in stercore."
Synonym: Ataenius rugiceps Dury, 1902: 153.
Type locality: "Cincinnati, O."
Genus Tesarius Rakovic, 1981
Tesarius Rakovic, 1981: 27 (Rakovic 1981a).
Type species: Phycochus sulcipennis Lea, 1904 by monotypy.
Tesarius caelatus (LeConte, 1857)
Original combination: Aegialia caelata LeConte, 1857: 42.
Type locality: "San Francisco."
Distribution: Nearctic; Palaearctic; Chile: VI Región del Bernardo O'Higgins (Skelley 2001).
Type genus: Scarabaeus Linnaeus, 1758: 345.
Tribe CANTHONINI Péringuey, 1901
Type genus: Canthon Hoffmannsegg, 1817: 38.
Genus Canthon (Canthon) Hoffmansegg, 1817
Canthon Hoffmannsegg, 1817: 38.
Type species: not verified.
Synonym: Coprobius Latreille, 1829: 535.
Type species: not verified.
Synonym: Coeloscelis Reiche, 1841: 213.
Type species: Coeloscelis coriaceus Reiche, 1841 by original designation.
Synonym: Paedhyboma Kolbe, 1893: 191.
Type species: not verified.
Synonym: Canthomoechus Pereira and Martínez, 1960: 166.
Type species: Canthon quadratus Blanchard, 1845 by original designation.
Canthon janthinus Blanchard, 1847
Original combination: Canthon janthinus Blanchard, 1845: 161 (Blanchard 1845b).
Type locality: "à la baie de San-Blas, dans l'île de Jabalis en Patagonie."
Distribution: Uruguay; Argentina: La Pampa, San Luis, Buenos Aires, Córdoba, Río Negro, Chubut (Blanchard 1845b, Martínez 1959b).
Genus Eudinopus Burmeister, 1840
Eudinopus Burmeister, 1840: [5].
Type species: Eudinopus ateuchoides Burmeister, 1840 by monotypy.
Eudinopus dytiscoides (Schreibers, 1802)
Original combination: Scarabaeus dytiscoides Schreibers, 1802: 191.
Type locality: "Brasilia."
Distribution: Argentina: Catamarca, La Pampa, San Juan, San Luis, La Rioja, Buenos Aires, Mendoza, Neuquén, Río Negro (Martínez 1959b).
Synonym: Eudinopus ateuchoides Burmeister, 1840: [6].
Type locality: "desertis Americae australis interioris, et in Paraguay inventa est."
Genus Megathopa Eschscholtz, 1822
Megathopa Eschscholtz, 1822: 34.
Type species: Megathopa villosa Eschscholtz, 1822 by monotypy.
Megathopa argentina Gillet, 1911
Original combination: Megathopa argentina Gillet, 1911: 315.
Type locality: "République Argentine: Cordobá..., Catamarca..., Chaco de Santiago."
Distribution: Uruguay; Argentina: Jujuy, Formosa, Catamarca, Santiago del Estero, Tucumán, Salta, La Pampa, San Luis, Córdoba, Entre Ríos, Buenos Aires, San Luis, La Rioja, Mendoza, Neuquén (Martínez 1950).
Synonym: Megathopa cuprea Gillet, 1911: 316.
Type locality: "Uruguay: Soriano."
Megathopa puncticollis puncticollis Blanchard, 1845
Original combination: Megathopa puncticollis Blanchard, 1845: 158 (Blanchard 1845b).
Type locality: "Patagonie."
Distribution: Bolivia; Paraguay; Argentina: Jujuy, Catamarca, Tucumán, Salta, Santiago del Estero, La Pampa, San Luis, Córdoba, Buenos Aires, La Rioja, Río Negro (Martínez 1950).
Megathopa villosa Eschscholtz, 1822
Original combination: Megathopa villosa Eschscholtz, 1822: 35.
Type locality: "Chili bei Conception."
Distribution: Argentina: Córdoba, Mendoza, Neuquén, Río Negro, Chubut, Santa Cruz; Chile: V Región deValparaíso, Región Metropolitana, VII-X Région (Gutiérrez 1940a, 1947; Martínez 1950).
Synonym: Megathopa porteri Gutiérrez, 1940: 99.
Type locality: "Viña del Mar... Los Angeles."
Megathopa violacea Blanchard, 1841
Original combination: Megathopa violacea Blanchard, 1841: plate 10.
Type locality: "Patagonie."
Distribution: Uruguay; Argentina: Catamarca, Misiones, La Pampa, San Luis, Córdoba, Buenos Aires, Río Negro (Martínez 1950).
Synonym: Megathopa picea Burmeister, 1874: 120.
Type locality: "Argentina."
Genus Scybalophagus Martínez, 1953
Scybalophagus Martínez, 1953: 3 (Martínez 1953b).
Type species: Scybalophagus patagonichus Martínez, 1953 by monotypy.
Scybalophagus lacordairei (Laporte, 1840)
Original combination: Hyboma lacordairei Laporte, 1840: 74.
Type locality: "Chili."
Distribution: Argentina: La Pampa, Buenos Aires, Río Negro (Martínez 1959b).
Scybalophagus patagonichus Martínez, 1953
Original combination: Scybalophagus patagonichus Martínez, 1953: 3 (Martínez 1953b).
Type locality: "Territorio Nacional de Chubut."
Distribution: Argentina: Neuquén, Río Negro, Chubut, Santa Cruz (Martínez 1959b).
Scybalophagus pilosus (Felsche, 1910)
Original combination: Canthon pilosus Felsche, 1910: 339.
Type locality: "Argentinien: Mendoza."
Distribution: Argentina: San Luis, Mendoza, Río Negro (Martínez 1959b).
Genus Tesserodoniella Vaz-de-Mello and Halffter, 2006
Tesserodoniella Vaz-de-Mello and Halffter, 2006: 60.
Type species: Tesserodoniella elguetai Vaz-de-Mello and Halffter, 2006 by original designation.
Tesserodoniella elguetai Vaz-de-Mello and Halffter, 2006
Original combination: Tesserodoniella elguetai Vaz-de-Mello and Halffter, 2006: 63.
Type locality: "Chile: Región Metropolitana de Santiago: Maipo, Rangue."
Distribution: Chile: Región Metropolitana (Vaz-de-Mello and Halffter 2006).
Tesserodoniella meridionalis Vaz-de-Mello and Halffter, 2006
Original combination: Tesserodoniella meridionalis Vaz-de-Mello and Halffter, 2006: 65.
Type locality: "Chile: VII Región del Maule: Constitución, Pantanillos."
Distribution: Chile: VII Región del Maule, VIII Región del Biobío (Vaz-de-Mello and Halffter 2006).
Tribe ATEUCHINI Laporte, 1840
Type genus: Ateuchus Weber, 1801: 10.
Genus Homocopris Burmeister, 1846
Homocopris Burmeister, 1846: [3].
Type species: Copris torulosus Eschscholtz, 1822 by monotypy.
Homocopris punctatissimus (Curtis, 1844)
Original combination: Copris punctatissima Curtis, 1844: 197.
Type locality: "Chiloe."
Distribution: Chile: X Région (Curtis 1844).
Synonym: Copris valdiviana Philippi, 1859: 661.
Type locality: "la provincia de Valdivia."
Homocopris torulosus (Eschscholtz, 1822)
Original combination: Copris torulosus Eschscholtz, 1822: 29.
Type locality: "Chili, Conception."
Distribution: Argentina: Neuquén, Chubut, Santa Cruz; Chile: VIII-XI Région (Gutiérrez 1940b, 1947).
Remark: The record of this species in Brasil is erroneous.
Synonym: Pinotus minor Luederwaldt, 1925: 68.
Type locality: "Chile e Estado do Espirito Santo."
Synonym: Pinotus dahli Landin, 1955: 5.
Type locality: "Chile: Isla Chiloé. S of Lechagua."
Tribe EUCRANIINI van Lansberge, 1874
Type genus: EucraniumBrullé, 1837: 289 (Audouin and Brullé 1837).
Genus Eucranium Brullé, 1837
Eucranium Brullé, 1837: 289 (Audouin and Brullé 1837).
Type species: Eucranium arachnoides Brullé, 1837 by monotypy.
Synonym: Anomiopsis Westwood, 1837: 13.
Type species: Anomiopsis dioscorides Westwood, 1837 by subsequent designation.
Synonym: Psammotrupes Guérin-Méneville, 1838: 45.
Type species: Psammotrupes dentifrons Guérin-Méneville, 1838 by monotypy.
Synonym: Cyclodema Laporte, 1840: 68.
Type species: Pachysoma lacordairei Laporte, 1840 by monotypy.
Eucranium arachnoides Brullé, 1837
Original combination: Eucranium arachnoides Brullé, 1837: 286 (Audouin and Brullé 1837).
Type locality: Not given.
Distribution: La Pampa, San Luis, Córdoba, Mendoza, Neuquén (Martínez 1959b).
Synonym: Anomiopsis dioscorides Westwood, 1837: 13.
Type locality: "Patagoniâ."
Synonym: Pachysoma lacordairei Laporte, 1840: 68.
Type locality: "Chili."
Eucranium dentifrons (Guérin-Méneville, 1838)
Original combination: Psammotrupes dentifrons Guérin-Méneville, 1838: 46.
Type locality: "Chili."
Distribution: Argentina: La Pampa, San Luis, Neuquén, Río Negro (Martínez 1959b).
Synonym: Anomiopsis aelinanus Blanchard, 1841: plate 10.
Type locality: "Patagonie."
Eucranium lepidum Burmeister, 1861
Original combination: Eucranium lepidum Burmeister, 1861: 61.
Type locality: "Bolivien" (erroneous).
Distribution: Argentina: Río Negro (Martínez 1959b).
Tribe PHANAEINI Kolbe, 1905
Type genus: Phanaeus MacLeay, 1819: 124.
Genus Sulcophanaeus D'Olsoufieff, 1924
Phanaeus (Sulcophanaeus) D'Olsoufieff, 1924: 23.
Type species: Scarabaeus sulcatus Drury, 1773.
Synonym: Eucopricus Gistel, 1857: 606.
Type species: Phanaeus columbi MacLeay, 1819.
Sulcophanaeus imperator (Chevrolat, 1844)
Original combination: Phanaeus imperator Chevrolat, 1830: 77 (Guérin-Méneville 1830).
Type locality: "le Brésil intérieur... le Tucuman et le Corrientes" (Guérin-Méneville 1844).
Distribution: Bolivia; Paraguay; Argentina: Formosa, Jujuy, Catamarca, Tucumán, Salta, Santiago del Estero, Santa Fe, Chaco, La Pampa, San Juan, San Luis, Corrientes, Córdoba, La Rioja, Buenos Aires, Mendoza (Martínez 1959b, Edmonds 2000).
Synonym: Phanaeus dimidiatus Solier, 1851: 63.
Type locality: "en la Repúblic" [Chile].
Tribe ONTHOPHAGINI Burmeister, 1846
Type genus: Onthophagus Latreille, 1802: 141.
Onthophagus (Digitonthophagus) Balthasar, 1959
Digitonthophagus Balthasar, 1959: 464.
Type species: not verified.
Onthophagus (Digitonthophagus) gazella (Fabricius, 1787)
Original combination: Scarabaeus gazella Fabricius, 1787: 377
Type locality: not verified.
Distribution: Easter Island (Ripa et al. 1995).
Genus Onthophagus (Onthophagus) Latreille, 1802
Onthophagus Latreille, 1802: 141.
Type species: Scarabaeus taurus Schreber, 1759 by monotypy.
Synonym: Chalcoderus Erichson, 1847: 763 (Erichson 1847b).
Type species: not verified.
Synonym: Monapus Erichson, 1847: 763 (Erichson 1847b).
Type species: not verified.
Synonym: Psilax Erichson, 1847: 764 (Erichson 1847b).
Type species: Onthophagus pronus Erichson, 1848 by monotypy.
Synonym: Gonocyphus van Lansberge, 1885: 382.
Type species: not verified.
Synonym: Tauronthophagus Shipp, 1895: 179.
Type species: Onthophagus rangifer Klug, 1855 by original designation.
Onthophagus hirculus Mannerheim, 1829
Original combination: Onthophagus hirculus Mannerheim, 1829: 39.
Type locality: "circa urbem Tijuco in stercoratis aridis" (Brazil).
Distribution: Brazil; Argentina: Buenos Aires, La Pampa, San Luis, Córdoba, San Juan, Santa Fe, Entre Ríos, Corrientes, Misiones, Chaco, Formosa, Santiago de Estero, Jujuy, Salta, Tucumán, Catamarca, La Rioja, Mendoza, Neuquén, Río Negro (Mannerheim 1829, Martínez 1959b).
Synonym: Onthophagus aeneus Blanchard, 1845: 183 (Blanchard 1845b).
Type locality: "la province de Valle Grande."
Synonym: Onthophagus brasiliensis Harold, 1859: 224 (replacement name for Onthophagus aeneus Blanchard, 1845).
Type locality: "la province de Valle Grande."
Tribe ONITINI Laporte, 1840
Type genus: Onitis Fabricius, 1798: 25
Genus Onitis Fabricius, 1798
Onitis Fabricius, 1798: 25
Type species: Scarabaeus sphinx Fabricius, 1775 by subsequent designation (Latreille 1810).
Onitis vanderkelleni van Lansberge, 1886
Original combination: Onitis vanderkelleni van Lansberge, 1886: 75.
Type locality: not verified.
Distribution: Africa (van Lansberge 1886), Easter Island (Ripa et al. 1995).
Type genus: Allidiostoma Arrow, 1940: 16.
Genus Allidiostoma Arrow, 1940
Allidiostoma Arrow, 1940: 16.
Type species: not verified.
Synonym: Idiostoma Arrow, 1904: 740.
Type species: not verified.
Allidiostoma bosqui Gutiérrez, 1946
Original combination: Allidiostoma bosqui Gutiérrez, 1946: 12.
Type locality: "El Canelo, Prv. De Santiago."
Distribution: Chile: V Región deValparaíso (Gutiérrez 1946).
Allidiostoma hirtum (Ohaus, 1910)
Original combination: Idiostoma hirta Ohaus, 1910: 173.
Type locality: "Argentina, Rio Chubut."
Distribution: Argentina: Chubut (Bruch 1911).
Allidiostoma landbecki (Philippi, 1873)
Original combination: Oryctes landbecki Philippi, 1873: 309.
Type locality: "Andibus de Talcaregue dictis."
Distribution: Argentina: Neuquén; Chile: VI Región del Bernardo O'Higgins (Philippi 1873, Roig-Juñent et al. 2005).
Synonym: Phyllognathus paulseni Fairmaire, 1885: CLXXXIX.
Type locality: "Concepcion."
Allidiostoma medon (Arrow, 1904)
Original combination: Idiostoma medon Arrow, 1904: 741.
Type locality: "Patagonia."
Distribution: "Patagonia" (Arrow 1904).
Allidiostoma porteri (Ruiz, 1924)
Original combination: Idiostoma porteri Ruiz, 1924: 100.
Type locality: "Valle del Lonquimay."
Distribution: Chile: IX Región de La Araucania (Ruiz 1924).
Allidiostoma rufum (Arrow, 1904)
Original combination: Idiostoma rufum Arrow, 1904: 741.
Type locality: "W. Patagonia, Valle del Lago Blanco."
Distribution: Argentina: Chubut; Chile: XII Región de Magallanes (Arrow 1904, Gutiérrez 1950).
Allidiostoma strobeli (Steinheil, 1874)
Original combination: Orphnus strobeli Steinheil, 1874: 556.
Type locality: "Bahia blanca."
Distribution: Argentina: Buenos Aires, "Patagonia" (Bruch 1911).
Type genus: Aclopus Erichson, 1835: 259.
Genus Aclopus Erichson, 1835
Aclopus Erichson, 1835: 259.
Type species: not verified.
Genus Neophaenognatha Allsopp, 1983
Neophaenognatha Allsopp, 1983: 209.
Type species: Phaenognatha jenseni Heller, 1908 by original designation.
Neophaenognatha minor (Bruch, 1910)
Original combination: Phaenognatha minor Bruch, 1910: 72.
Type locality: "Gob. Del Rio Negro."
Distribution: Argentina: Río Negro (Bruch 1910).
Type genus: Melolontha Fabricius, 1775: 31.
Type genus: Lichnia Erichson, 1835: 269.
Genus Arctodium Burmeister, 1844
Arctodium Burmeister, 1844: 9.
Type species: Cratoscelis vulpina Erichson, 1835 by subsequent designation (Chapin 1938).
Remark: Although Arctodium was first used by Dejean (1833), Burmeister (1844) should be credited with authorship. Burmeister (1844) listed Arctodium as an invalid synonym of Cratoscelis Erichson but later use as a valid genus by Chapin (1938) makes Arctodium available under Article 11.6.1 of the Code with Burmeister (1844) as the author.
Synonym: Cratoscelis Erichson, 1835: 267.
Type species: Cratoscelis vulpina Erichson, 1835 by subsequent designation (Chapin 1938).
Arctodium discolor (Erichson, 1835)
Original combination: Cratoscelis discolor Erichson, 1835: 269.
Type locality: "Chile."
Distribution: Chile: VIII Región del Biobío, IX Región de la Araucania (Hawkins 2006).
Arctodium mahdii Hawkins, 2006
Original combination: Arctodium mahdii Hawkins, 2006: 37.
Type locality: "Prov. Linares, Cord. Parral, Fdo. Malcho."
Distribution: Chile: VII-IX Región (Hawkins 2006).
Arctodium planum (Blanchard, 1850)
Original combination: Cratoscelis plana Blanchard, 1850: 52.
Type locality: "Chile."
Distribution: Chile: V Región de Valparaíso, VI Región del O'Higgins, Región Metropolitana (Hawkins 2006).
Synonym: Cratoscelis canicapilla Philippi and Philippi, 1864: 326.
Type locality: "Valparaiso."
Arctodium vulpinum (Erichson, 1835)
Original combination: Cratoscelis vulpina Erichson, 1835: 269.
Type locality: "Chile."
Distribution: Chile: IV-IX Región, Región Metropolitana (Hawkins 2006).
Synonym: Cratoscelis aterrima Blanchard, 1850: 53.
Type locality: "Chili."
Synonym: Cratoscelis gayana Blanchard, 1850: 53.
Type locality: "Chile."
Synonym: Cratoscelis villosa Blanchard, 1850: 53.
Type locality: "Chile."
Synonym: Cratoscelis striolata Redtenbacher, 1868: 61.
Type locality: "Chili."
Genus Lichnia Erichson, 1835
Lichnia Erichson, 1835: 269.
Type species: Lichnia limbata Erichson, 1835 by monotypy.
Remark: Gutiérrez (1943) attempted to create the subgenus Neolichnia within this genus but since no type species was designated for this name in the original publication it is a nomen nudum.
Synonym: Dasychaeta Erichson, 1847:104 (Erichson 1847a).
Type species: Dasychaeta lateralis Erichson, by monotypy.
Lichnia gallardoi Gutiérrez, 1943
Original combination: Lichnia gallardoi Gutiérrez, 1943: 151.
Type locality: "Huetulame, (provincia de Coquimbo)."
Distribution: Chile: IV Región de Coquimbo (Hawkins 2006).
Synonym: Lichnia porteri Gutiérrez, 1943: 152.
Type locality: "Hurtado, (Ovalle)... y Huetulame."
Lichnia limbata Erichson, 1835
Original combination: Lichnia limbata Erichson, 1835: 270.
Type locality: "Chile."
Distribution: Chile: V-VII Región, Región Metropolitana (Hawkins 2006).
Synonym: Dasychaeta lateralis Erichson, 1847:104 (Erichson 1847a).
Type locality: "Republica Peruana" (erroneous).
Synonym: Lichnia immarginata Blanchard, 1850: 52.
Type locality: "Chili."
Type genus: Serica MacLeay, 1819: 146.
Genus Athlia Erichson, 1835
Athlia Erichson, 1835: 266.
Type species: Athlia rustica Erichson, 1835 by monotypy.
Remark: Key to species in Saylor, 1946b: 21; Gutiérrez, 1949: 13; Martínez, 1967: 337; 1974: 352.
Synonym: Rivera Germain, 1903: 392 (Germain 1903a).
Type species: Athlia plebeja Burmeister, 1855 by monotypy.
Subgenus: Dihymenonyx Gutiérrez, 1949: 16.
Type species: Dihymenonyx herrerai Gutiérrez, 1949 by monotypy.
Athlia bruchi Moser, 1924
Original combination: Athlia bruchi Moser, 1924a: 121.
Type locality: "Argentina: Cordoba."
Distribution: Argentina: Córdoba, San Luis, Buenos Aires, La Pampa, Neuquén, Río Negro (Gutiérrez 1949, Martínez 1967).
Athlia giaii Martínez, 1959
Original combination: Athlia giaii Martínez, 1959a: 23.
Type locality: "Neuquén, Piedra del Aguila."
Distribution: Argentina: Neuquén, Río Negro (Martínez 1959a).
Athlia herrerai (Gutiérrez, 1949)
Original combination: Dihymenonyx herrerai Gutiérrez, 1949: 16.
Type locality: "Chile: Territorio de Magallanes, Punta Arenas."
Distribution: Chile: Región XII de Magallanes y la Antartica Chilena (Gutiérrez 1949).
Athlia parvissima Saylor, 1946
Original combination: Athlia parvissima Saylor, 1946b: 24.
Type locality: "Zapala, Neuquen, Argentina."
Distribution: Argentina: Neuquén, Río Negro (Martínez 1959a).
Athlia plebeja Burmeister, 1855
Original combination: Athlia plebeja Burmeister, 1855: 125.
Type locality: "Chili."
Distribution: Chile: V Región de Valparaíso, VIII Región del Biobío (Martínez 1967).
Synonym: Athlia rivera Saylor, 1946b: 23.
Type locality: "Chile, Concepcion."
Athlia problematica Gutiérrez, 1950
Original combination: Athlia problematica Gutiérrez, 1950a: 270.
Type locality: "Chile: Prov. de Valparaíso, Concón."
Distribution: Chile: V Región de Valparaíso (Gutiérrez 1949), VI Región del Bernardo O'Higgins (Martínez 1967).
Athlia rotundata Gutiérrez, 1949
Original combination: Athlia rotundata Gutiérrez, 1949: 12.
Type locality: Chile: "Prov. Valparaíso, Concón."
Distribution: Chile: V Región de Valparaíso (Gutiérrez 1949).
Athlia rustica Erichson, 1835
Original combination: Athlia rustica Erichson, 1835: 267.
Type locality: "Chile."
Distribution: Chile: V-VII Región (Martínez 1967).
Type genus: Sericoides Guérin-Méneville, 1840: 301.
Sericoides Guérin-Méneville, 1839: 301.
Type species: Melolontha glacialis Fabricius, 1775 by subsequent designation (Evans 2003).
Synonym: Macrosoma Hope, 1837: 109 (preoccupied).
Type species: Melolontha glacialis Fabricius, 1775 by original designation.
Synonym: Listronyx Guérin-Méneville, 1839: 302.
Type species: Listronyx nigriceps Guérin-Méneville, 1839 by monotypy.
Synonym: Accia Curtis, 1844: 199.
Type species: Accia lucida Curtis, 1844 by monotypy.
Synonym: Maypa Blanchard, 1850: 115.
Type species: Maypa chlorosticta Blanchard, 1850 by monotypy.
Synonym: Apterodema Fairmaire, 1884: 490.
Type species: Apterodema acuticollis Fairmaire, 1884 by monotypy.
Synonym: Paralistronyx Brenske, 1906: 30.
Type species: Listronyx livida Germain, 1863 by original designation.
Synonym: Paramaypa Brenske, 1906: 30.
Type species: Maypa osornoana Brenske, 1906 by original designation.
Sericoides acuticollis (Fairmaire, 1884)
Original combination: Apterodema acuticollis Fairmaire, 1884: 491
Type locality: "Punta-Arena."
Distribution: Argentina: Tierra del Fuego, Chile: XII Región de Magallanes (Gutiérrez 1947, Martínez 1972b).
Sericoides andina (Germain, 1863)
Original combination: Maypa andina Germain, 1863: 726 (in Fairmaire and Germain 1863).
Type locality: "de hauteur dans les Cordillères de Chillan."
Distribution: Chile: VIII Región del Biobío (Fairmaire and Germain 1863).
Sericoides antarctica (Brenske, 1900)
Original combination: Listronyx antarcticus Brenske, 1900: 109.
Type locality: "Canal du Beagle, Forêt de Lapataïa."
Distribution: Argentina: Tierra del Fuego; Chile: XII Región de Magallanes (Gutiérrez 1947).
Sericoides argentinensis Martínez, 1971
Original combination: Sericoides (Listronyx) argentinensis Martínez, 1971: 115.
Type locality: "Argentina, provincia del Neuquén, estancia Fortín Chacabuco, situada en las proximidades del río Limay y del lago Nahuel Huapi."
Distribution: Argentina: Neuquén, Río Negro (Martínez 1971).
Sericoides atra (Solier, 1851)
Original combination: Maypa atra Solier, 1851: 108
Type locality: "Tambien se halla en el sur de la República" (Chile).
Distribution: Chile (Solier 1851).
Sericoides castanea Guérin-Méneville, 1839
Original combination: Sericoides castanea Guérin-Méneville, 1839: 302.
Type locality: "Chili."
Distribution: Argentina: "Patagonia;" Chile: VIII Región del Biobío, X Región de Los Lagos (Bruch 1911, Gutiérrez 1947).
Sericoides chilena (Germain, 1863)
Original combination: Maypa chilena Germain, 1863: 736 (in Fairmaire and Germain 1863).
Type locality: "provient des environs de Santiago."
Distribution: Chile: Región Metropolitana (Fairmaire and Germain 1863).
Sericoides chiliensis Blanchard, 1850
Original combination: Sericoides chiliensis Blanchard, 1850: 114.
Type locality: "Chili."
Distribution: Chile (Blanchard 1850).
Sericoides chlorosticta (Blanchard, 1850)
Original combination: Maypa chlorosticta Blanchard, 1850: 115.
Type locality: "Chili."
Distribution: Chile: VII Región del Biobío, X Región de Los Lagos (Philippi 1861, Gutiérrez 1949).
Junior synonym: Sericoides similis Philippi, 1861: 737
Type locality: "Valdivia."
Sericoides comata (Germain, 1863)
Original combination: Maypa comata Germain, 1863: 739 (in Fairmaire and Germain 1863).
Type locality: "Valdivia."
Distribution: Chile: X Región de Los Lagos (Fairmaire and Germain 1863).
Sericoides convexa (Germain, 1863)
Original combination: Maypa convexa Germain, 1863: 729 (in Fairmaire and Germain 1863).
Type locality: "Valdivia."
Distribution: Chile: X Región de Los Lagos (Fairmaire and Germain 1863).
Sericoides delicatula (Germain, 1863)
Original combination: Maypa delicatula Germain, 1863: 731 (in Fairmaire and Germain 1863).
Type locality: "Chiloë."
Distribution: Chile: X Región de Los Lagos (Gutiérrez 1949).
Sericoides dubia (Germain, 1863)
Original combination: Maypa dubia Germain, 1863: 738 (in Fairmaire and Germain 1863).
Type locality: "forêts subandines de Chillan."
Distribution: Chile: VIII Región del Biobío (Fairmaire and Germain 1863).
Sericoides faminaei (Blanchard, 1850)
Original combination: Listronyx faminaei Blanchard, 1850: 114.
Type locality: "Port-Famine."
Distribution: Argentina: Tierra del Fuego; Chile: XI Región Aisen, XII Región de Magallanes (Gutiérrez 1947).
Sericoides frigida (Germain, 1863)
Original combination: Listronyx frigida Germain, 1863: 743 (in Fairmaire and Germain 1863).
Type locality: "Chili."
Distribution: Chile (Fairmaire and Germain 1863).
Sericoides germaini Dalla Torre, 1912
Original combination: Sericoides germaini Dalla Torre, 1912b: 131 (for Germain, 1862: 741).
Type locality: "Cordilières de Chillan; 2,000 m."
Distribution: Argentina: Chubut; Chile: VIII Región del Biobío (Bruch 1911, Gutiérrez 1949).
Synonym: Listronyx castanea Germain, 1863: 741 (in Fairmaire and Germain 1863) (not Guérin-Méneville, 1839: 302).
Type locality: "Cordilières de Chillan; 2,000 m."
Sericoides glacialis (Fabricius, 1775)
Original combination: Melolontha glaciale Fabricius, 1775: 35.
Type locality: "Tierra del Fuego."
Distribution: Argentina: Chubut, Tierra del Fuego; Chile: X Región de Los Lagos, XII Región de Magallanes (Bruch 1911, Gutiérrez 1947).
Synonym: Sericoides reichei Guérin-Méneville, 1839: 301.
Type locality: not given.
Synonym: Sericoides atricapilla Curtis, 1844: 199.
Type locality: "Straits of Magellan."
Sericoides hirsuta (Brenske, 1900)
Original combination: Listronyx hirsutus Brenske, 1900: 110
Type locality: "Cenio del Ultima Esperanza Magellanes."
Distribution: Argentina: Tierra del Fuego; Chile: XII Región de Magallanes (Gutiérrez 1947).
Sericoides intermedia (Philippi, 1864)
Original combination: Maypa intermedia Philippi 1864: 452.
Type locality: "Andes prov. Santiago."
Distribution: Chile: VI Región del Bernardo O'Higgins (Philippi 1864).
Sericoides lineolata (Germain, 1863)
Original combination: Maypa lineolata Germain, 1863: 732 (in Fairmaire and Germain 1863).
Type locality: "Valdivia, Chillan."
Distribution: Chile: VIII Región del Biobío, X Región de Los Lagos (Gutiérrez 1949).
Sericoides livida (Germain, 1863)
Original combination: Listronyx livida Germain, 1863: 745 (in Fairmaire and Germain 1863).
Type locality: "Chili."
Distribution: Argentina: Neuquén, Chubut, Tierra del Fuego; Chile: VIII Región del Biobío (Martínez 1959a, 1972b).
Distribution: Argentina: Tierra del Fuego; Chile: XI Región Aisen, XII Región de Magallanes (Bruch 1911, Gutiérrez 1947).
Synonym: Listronyx obscura Philippi, 1864: 453.
Type locality: "Corral."
Remark: junior secondary junior homonym of Sericoides obscura (Germain, 1863).
Synonym: Macrosoma (Paramaypa) osornoana Brenske, 1906: 30.
Type locality: not given.
Synonym: Sericoides (Listronyx) aenigmaticus Martínez, 1959: 27 (Martínez 1959a) (for Listronyx obscura Philippi 1864: 453).
Type locality: "Corral."
Remark: replacement name for Listronyx obscura Philippi, 1864.
Sericoides lloydi Martínez, 1965
Original combination: Sericoides lloydi Martínez, 1965: 89.
Type locality: "Argentina, provincia de Río Negro, Bariloche."
Distribution: Argentina: Río Negro (Martínez 1965).
Sericoides longipes (Germain, 1863)
Original combination: Maypa longipes Germain, 1863: 727 (in Fairmaire and Germain 1863).
Type locality: "Forêts subandines de Chillan."
Distribution: Chile: VIII Región del Biobío (Fairmaire and Germain 1863).
Sericoides lucida (Curtis, 1844)
Original combination: Accia lucida Curtis, 1844: 200
Type locality: "Port St. Elena."
Distribution: Argentina: Tierra del Fuego; Chile: XI Región Aisen, XII Región de Magallanes (Bruch 1911, Gutiérrez 1947).
Sericoides monticola (Germain, 1863)
Original combination: Maypa monticola Germain, 1863: 736 (in Fairmaire and Germain 1863)
Type locality: "montagnes du Chili central, à 1,800 m."
Distribution: Chile
Sericoides multicolor Martínez, 1956
Original combination: Sericoides multicolor Martínez, 1956: 45.
Type locality: "Argentina, Tierra del Fuego, Bosque de Lapataia."
Distribution: Argentina: Tierra del Fuego; Chile: XII Región de Magallanes (Martínez 1972b).
Sericoides nitida Philippi and Philippi, 1864
Original combination: Sericoides nitida Philippi and Philippi, 1864: 325.
Type locality: "Corral."
Distribution: Chile: X Región de Los Lagos (Philippi and Philippi 1864).
Sericoides nossi Martínez, 1972
Original combination: Sericoides nossi Martínez 1972b: 50.
Type locality: "Argentina, provincia de Neuquén, Rincón Chico (estancias San Ramón, Tehuel Malal y Jones)."
Distribution: Argentina: Neuquén (Martínez 1972b).
Sericoides obesa (Germain, 1863)
Original combination: Maypa obesa Germain, 1863: 728 (in Fairmaire and Germain 1863).
Type locality: "Forêts subandines de Chillan."
Distribution: Chile: VIII Región del Biobío (Gutiérrez 1949).
Sericoides obscura (Germain, 1863)
Original combination: Maypa obscura Germain, 1863: 728 (in Fairmaire and Germain 1863).
Type locality: "Forêts subandines de Chillan."
Distribution: Argentina: Neuquén; Chile: X Región de Los Lagos, XI Región Aisen (Gutiérrez 1947).
Sericoides olivacea (Germain, 1863)
Original combination: Maypa olivacea Germain, 1863: 735 (in Fairmaire and Germain 1863).
Type locality: "Forêts subandines de Chillan."
Distribution: Chile: VIII Región del Biobío (Fairmaire and Germain 1863).
Sericoides opacipennis (Germain, 1863)
Original combination: Maypa opacipennis Germain, 1863: 738 (in Fairmaire and Germain 1863).
Type locality: "Santiago; sous une pierre."
Distribution: Argentina: Neuquén; Chile: Región Metropolitana, X Región de Los Lagos (Gutiérrez 1947).
Sericoides pallida (Germain, 1863)
Original combination: Listronyx pallida Germain, 1863: 744 (in Fairmaire and Germain 1863).
Type locality: "à 2,500 mètres, dans les Cordilières de Chillan."
Distribution: Chile: VIII Región del Biobío (Fairmaire and Germain 1863).
Sericoides palpalis (Germain, 1863)
Original combination: Maypa palpalis Germain, 1863: 725 (in Fairmaire and Germain 1863)
Type locality: "Chiloë"
Distribution: Chile: X Región de Los Lagos (Fairmaire and Germain 1863)
Sericoides philippiana Dalla Torre, 1912
Original combination: Sericoides philippiana Dalla Torre, 1912b: 132 (for Philippi, 1864: 451).
Type locality: "In andibus prov. Santiago sub Chuquiraga oppositifolia (Yerba blanca) invenit orn."
Distribution: Chile: Región Metropolitana (Philippi 1864).
Synonym: Maipa andina Philippi, 1864: 451 (not Germain, 1863: 726).
Type locality: "In andibus prov. Santiago sub Chuquiraga oppositifolia (Yerba blanca) invenit orn."
Sericoides picea (Kolbe, 1907)
Original combination: Accia picea Kolbe, 1907: 111.
Type locality: "Süd-Patagonien, Punta Arenas,... Süd-Feuerland, Uschuaia,... Isla Navarin, Puerto Toro."
Distribution: Argentina Tierra del Fuego; Chile: XI Región Aisen, XII Región de Magallanes (Bruch 1911, Gutiérrez 1947).
Sericoides piligera (Germain, 1863)
Original combination: Maypa piligera Germain, 1863: 731 (in Fairmaire and Germain 1863).
Type locality: "à environ 2,000 m. dans les Cordilières de Chillan."
Distribution: Chile: VIII Región del Biobío (Fairmaire and Germain 1863).
Sericoides pubescens (Germain, 1863)
Original combination: Maypa pubescens Germain, 1863: 733 (in Fairmaire and Germain 1863).
Type locality: "Forêts subandines de Chillan."
Distribution: Chile: VIII Región del Biobío (Fairmaire and Germain 1863).
Sericoides punctata (Solier, 1851)
Original combination: Maypa punctata Solier, 1851: 107.
Type locality: "Chile."
Distribution: Chile (Solier 1851).
Sericoides rechencqui Martínez, 1972
Original combination: Sericoides rechencqui Martínez 1972b: 45.
Type locality: "Argentina, provincia de Neuquén, Estancia Fortín Chacabuco, en las proximidades del río Limay y del lago Nahuel Huapí."
Distribution: Argentina: Neuquén (Martínez 1972b).
Sericoides rufeola (Solier, 1851)
Original combination: Maypa rufeola Solier, 1851: 108.
Type locality: "Chile."
Distribution: Chile (Solier 1851).
Sericoides rufocastanea (Germain, 1863)
Original combination: Maypa rufocastanea Germain, 1863: 737 (in Fairmaire and Germain 1863).
Type locality: "Santiago."
Distribution: Chile: Región Metropolitana (Fairmaire and Germain 1863).
Sericoides rugosula (Germain, 1863)
Original combination: Maypa rugosula Germain, 1863: 733 (in Fairmaire and Germain 1863).
Type locality: "Forêts subandines de Chillan, prise le soir au vol."
Distribution: Chile: VIII Región del Biobío (Fairmaire and Germain 1863).
Synonym: Maypa cuprea Philippi, 1864: 450.
Type locality: "Valdivia."
Sericoides sinuaticollis (Germain, 1863)
Original combination: Maypa sinuaticollis Germain, 1863: 737 (in Fairmaire and Germain 1863).
Type locality: "Valdivia."
Distribution: Chile: X Región de Los Lagos (Fairmaire and Germain 1863).
Sericoides striata (Fabricius, 1775)
Original combination: Melolontha striata Fabricius, 1775: 35.
Type locality: "Tierra del Fuego."
Distribution: Chile: XII Región de Magallanes (Fabricius 1775).
Synonym: Maypa striata Philippi, 1864: 451.
Type locality: "In andibus prov. Santiago ocurrit."
Sericoides subcostata (Germain, 1863)
Original combination: Maypa subcostata Germain, 1863: 729 (in Fairmaire and Germain 1863).
Type locality: "Valdivia."
Distribution: Chile: X Región de Los Lagos (Fairmaire and Germain 1863).
Synonym: Maypa opaca Philippi and Philippi, 1864: 324.
Type locality: "Corral."
Sericoides sulcatopunctata (Germain, 1863)
Original combination: Listronyx sulcato-punctata Germain, 1863: 740 (in Fairmaire and Germain 1863).
Type locality: "dans les vallées subandines de la province de Santiago."
Distribution: Chile: Región Metropolitana (Fairmaire and Germain 1863).
Sericoides sylvatica (Germain, 1863)
Original combination: Maypa sylvatica Germain, 1863: 730 (in Fairmaire and Germain 1863).
Type locality: "forêts subandines de Chillan; sous les écorces."
Distribution: Argentina: Neuquén, Río Negro; Chile: VIII Región del Biobío (Bruch 1911, Gutiérrez 1947).
Sericoides testacea (Fabricius, 1775)
Original combination: Melolontha testacea Fabricius, 1775: 35.
Type locality: "Tierra del Fuego."
Distribution: Argentina: Chubut, Tierra del Fuego; Chile: X Región de Los Lagos, XII Región de Magallanes (Bruch 1911, Gutiérrez 1947).
Synonym: Melolontha fusca Gmelin, 1790: 1569.
Type locality: "terra del fuego."
Synonym: Listronyx nigriceps Guérin-Méneville, 1839: 302.
Type locality: "Terre-de-Feu."
Synonym: Listronyx melanocephala Blanchard, 1846: plate 8.
Type locality: "la Terre-de-Feu, n'a été retrouvé que depuis peu d'années au port Famine, dans le détroit de Magellan" (Blanchard 1854).
Sericoides variegata (Germain, 1863)
Original combination: Listronyx variegata Germain, 1863: 742 (in Fairmaire and Germain 1863) (for Solier, 1851: 109)
Type locality: "Se encuentra tambien en el sur de la República" (Chile).
Distribution: Chile (Solier 1851)
Junior synonym: Maypa chlorosticta Solier, 1851: 109 (not Blanchard, 1850: 115)
Type locality: "Se encuentra tambien en el sur de la República" (Chile).
Sericoides variolosa (Solier, 1851)
Original combination: Maypa variolosa Solier, 1851: 109.
Type locality: "Chile."
Distribution: Chile (Solier 1851).
Synonym: Sericoides rugulosa Philippi 1861: 736.
Type locality: "Valdivia."
Sericoides vestita (Germain, 1863)
Original combination: Listronyx vestita Germain, 1863: 744 (in Fairmaire and Germain 1863).
Type locality: "à 2,000 m., dans les Cordilières de Chillan."
Distribution: Argentina: Tierra del Fuego; Chile: VIII Región del Biobío, XII Región de Magallanes (Gutiérrez 1947, 1949).
Sericoides viridis (Solier, 1851)
Original combination: Maypa viridis Solier, 1851: 107.
Type locality: "Concepcion y en la Araucania."
Distribution: Argentina: Neuquén, Río Negro; Chile: VIII Región del Biobío, IX Región de la Araucania, XI Región Aisen (Solier 1851; Bruch 1911; Gutiérrez 1947, 1949).
Genus Ulata Saylor, 1945
Ulata Saylor, 1945c: 441.
Type species: Ulata argentina Saylor, by original designation.
Ulata argentina Saylor, 1945
Original combination: Ulata argentina Saylor, 1945c: 442.
Type locality: "Bariloche, Rio Negro, Argentina."
Distribution: Argentina: Río Negro (Saylor 1945c); Chile: VIII Región del Biobío (Gutiérrez 1949).
Type genus: Liparetrus Guérin-Méneville, 1831: 3.
Genus Aplodema Blanchard, 1850
Aplodema Blanchard, 1850: 115.
Type species Brachyphylla magellanica Blanchard, 1842 by subsequent designation (Evans 2003).
Junior synonym: Haplodema Harold, 1869a: 1136 (unjustified emendation)
Type species Brachyphylla magellanica Blanchard, 1842 by subsequent designation (Evans 2003).
Aplodema magellanica (Blanchard, 1842)
Original combination: Brachyphylla magellanica Blanchard, 1842: plate 8.
Type locality: "port Famine, détroit de Magellan."
Distribution: Argentina: Tierra del Fuego; Chile: XII Región de Magallanes (Gutiérrez1947).
Type genus: Diplotaxis Kirby, 1837: 129.
Liogenys Guérin-Méneville, 1831b: plate 3.
Type species: Melolontha palpalis Eschscholtz, 1822 by monotypy.
Remark: Key to species in Gutiérrez, 1951: 132; Frey, 1969a: 38.
Synonym: Amphicrania Dejean, 1833: 163.
Type species: Melolontha palpalis Eschscholtz, 1822 by monotypy.
Liogenys flaveola Moser, 1924
Original combination: Liogenys flaveolus Moser, 1924: 122 (Moser 1924a).
Type locality: "Argentina."
Distribution: Argentina (Moser 1924a).
Liogenys flavida Moser, 1918
Original combination: Liogenys flavidus Moser, 1918: 106 (Moser 1918b).
Type locality: "Patagonien."
Distribution: Argentina: "Patagonia" (Moser 1918b).
Liogenys grandis Philippi and Philippi, 1864
Original combination: Liogenys grandis Philippi and Philippi, 1864: 322.
Type locality: "Illapel, Santa Rosa."
Distribution: Chile: IV Región de Coquimbo, VI Región del Bernardo O'Higgins (Philippi and Philippi 1864, Gutiérrez 1949).
Liogenys hirta Gutiérrez, 1951
Original combination: Liogenys hirtus Gutiérrez, 1951: 139.
Type locality: "Coquimbo: Estación Pelícana, ramal a Rivadavia."
Distribution: Chile: IV Región de Coquimbo (Gutiérrez 1951).
Liogenys kadleci Frey, 1970
Original combination: Liogenys kadleci Frey 1970: 281.
Type locality: "Argentinien, Villa Regina, Rio Negro."
Distribution: Argentina: Río Negro (Frey 1970).
Liogenys kuntzeni Moser, 1921
Original combination: Liogenys kuntzeni Moser, 1921b: 139.
Type locality: "Chile."
Liogenys morio Burmeister, 1855
Original combination: Liogenys morio Burmeister, 1855: 16.
Type locality: "Mittel Amerika."
Distribution: Argentina: Chaco, Buenos Aires, Córdoba, "Patagonia" (Bruch 1911).
Liogenys obesula Gutiérrez, 1951
Original combination: Liogenys obesulus Gutiérrez, 1951: 135.
Type locality: "Curicó: Pichibudis (Curicó marítimo)."
Distribution: Chile: VII Región del Maule (Gutiérrez 1951).
Liogenys obscura Blanchard, 1851
Original combination: Liogenys obscurus Blanchard, 1851: 167.
Type locality: "Patagonie (Rio-Negro)."
Distribution: Argentina: Buenos Aires, San Luis, Río Negro (Blanchard 1851, Bruch 1911).
Synonym: Liogenys bruchi Moser, 1924a: 122.
Type locality: "Argentina: Alta Gracia La Granja, Sierras de Cordoba."
Liogenys pallens Blanchard, 1851
Original combination: Liogenys pallens Blanchard, 1851: 167.
Type locality: "Montevideo."
Distribution: Uruguay; Argentina: Chubut (Bruch 1911, Evans 2003).
Liogenys palpalis (Eschscholtz, 1822)
Original combination: Melolontha palpalis Eschscholtz, 1822: 17.
Type locality: "Chili, Conception."
Distribution: Chile: IV Región de Coquimbo, VI-VIII Región (Gutiérrez 1949).
Synonym: Liogenys castaneus Guérin-Méneville, 1831b: plate 3.
Type locality: "Conception, au Chili."
Synonym: Liogenys reichei Germain, 1903: 51 (Germain 1903b).
Type locality: "La Mocha."
Liogenys penai Gutiérrez, 1951
Original combination: Liogenys peñai Gutiérrez, 1951: 141.
Type locality: "Coquimbo Rivadavia."
Distribution: Chile: IV Región de Coquimbo (Gutiérrez 1951).
Liogenys wagenknechti Gutiérrez, 1951
Original combination: Liogenys wagenknechti Gutiérrez, 1951: 137.
Type locality: "Coquimbo: San Julián, Costa de Ovalle; Llanos de Chimba; Llanos de Barraza."
Distribution: Chile: IV Región de Coquimbo (Gutiérrez 1951).
Liogenys xanthocera Harold, 1869
Original combination: Liogenys xanthocerus Harold, 1869: 123 (Harold 1869c) (replacement name for Liogenys pallidicornis Blanchard, 1851: 168).
Type locality: "Patagonie (baie de San-Blas)."
Distribution: Argentina: Chubut (Harold 1869c).
Synonym: Liogenys pallidicornis Blanchard, 1851: 168 (not Liogenys pallidicornis Blanchard, 1851: 167).
Type locality: "Patagonie (baie de San-Blas)."
Pachrodema Blanchard, 1851: 168.
Type species: Pachrodema castanea Blanchard, 1851 by subsequent designation (Evans 2003).
Pachrodema castanea Blanchard, 1851
Original combination: Pachrodema castanea Blanchard, 1851: 168.
Type locality: "Patagonie (baie de San-Blas)."
Distribution: Argentina: Chubut; Chile: IV Región de Coquimbo, VI-IX Región (Blanchard 1851, Gutiérrez 1951).
Pachrodema cuprascens Burmeister, 1855
Original combination: Pachrodema cuprascens Burmeister, 1855: 260.
Type locality: "Patagonien."
Distribution: Argentina: "Patagonia" (Burmeister 1855).
Pachrodema lucida Burmeister, 1881
Original combination: Pachrodema lucida Burmeister, 1881: 100 (in Berg 1881).
Type locality: "Patagonia" and "Salinas Chicas."
Distribution: Argentina: Río Negro (Berg 1881).
Pachrodema picea Nonfried, 1894
Original combination: Pachrodema picea Nonfried, 1894: 115.
Type locality: "Santa Cruz, Patagonia."
Distribution: Argentina: Santa Cruz (Nonfried 1894).
Pachrodema substriata Blanchard, 1851
Original combination: Pachrodema substriata Blanchard, 1851: 169.
Type locality: "Patagonie (baie de San-Blas)."
Distribution: Argentina: Chubut (Blanchard 1851).
Genus Pacuvia Curtis, 1844
Pacuvia Curtis, 1844: 199.
Type species: Pacuvia castanea Curtis, by monotypy.
Remark: Key to species in Gutiérrez, 1951: 143.
Pacuvia castanea Curtis, 1844
Original combination: Pacuvia castanea Curtis, 1844: 199.
Type locality: "Valparaiso."
Distribution: Chile: IV-IX Región (Curtis 1845, Gutiérrez 1951).
Synonym: Liogenys gayana Solier, 1851: 100.
Type locality: "Santiago, Santa Rosa, Illapel, etc."
Pacuvia philippiana Gutiérrez, 1951
Original combination: Pacuvia philippiana Gutiérrez, 1951: 144.
Type locality: "Coquimbo: Fray Jorge."
Distribution: Chile: IV Región de Coquimbo (Gutiérrez 1951).
Type genus: Pachydema Laporte, 1840: 130.
Genus Diaphylla Erichson, 1847
Diaphylla Erichson, 1847: 102 (Erichson 1847a).
Type species: Diaphylla hispida Erichson, 1847 by monotypy.
Diaphylla granulata Philippi, 1864
Original combination: Diaphylla granulata Philippi, 1864: 439.
Type locality: "E prov. Valdivia attuli."
Distribution: Argentina: Neuquén (Ruiz-Manzanos 2006); Chile: X Región de Los Lagos (Philippi 1864).
Diaphylla hispida Erichson, 1847
Original combination: Diaphylla hispida Erichson, 1847: 102 (Erichson 1847a).
Type locality: "Republica Peruana."
Distribution: Chile: V Región de Valparaíso (Gutiérrez 1949).
Diaphylla lampropyga Philippi, 1864
Original combination: Diaphylla lampropyga Philippi, 1864: 438.
Type locality: "Corral in prov. Valdivia."
Distribution: Chile: X Región de Los Lagos (Philippi 1864).
Diaphylla luctuosa Philippi and Philippi, 1864
Original combination: Diaphylla luctuosa Philippi and Philippi, 1864: 323.
Type locality: "Valdivia."
Distribution: Chile: X Región de Los Lagos (Philippi and Philippi 1864).
Diaphylla ornata Philippi, 1864
Original combination: Diaphylla ornata Philippi, 1864: 439.
Type locality: "Prope Corral in prov. Valdivia reperitur."
Distribution: Chile: X Región de Los Lagos (Philippi 1864).
Faargia Martínez, 1982: 89.
Type species: Myloxena gentilii Martínez, by original designation.
Faargia gentillii (Martínez, 1975)
Original combination: Myloxena gentillii Martínez, 1975: 239.
Type locality: "la provincia de Neuquén, Zapala, Bardita."
Distributon: Argentina: Neuquén (Martínez 1975).
Genus Leuretra Erichson, 1847
Leuretra Erichson, 1847: 100 (Erichson 1847a).
Type species: Leuretra pectoralis Erichson, 1847 by monotypy.
Leuretra pectoralis Erichson, 1847
Original combination: Leuretra pectoralis Erichson, 1847: 101 (Erichson 1847a).
Type locality: "Republica Peruana."
Distribution: Peru; Chile (?) (Martínez 1975).
Genus Myloxena Burmeister, 1881
Myloxena Burmeister, 1881: plate 2 (in Berg 1881).
Type species: Myloxena vestita Burmeister, 1881 by monotypy.
Myloxena bruchiana Ohaus, 1909
Original combination: Myloxena bruchiana Ohaus, 1909: 440 (Ohaus 1909a).
Type locality: "Argentina, Gob. del Chubut."
Distribution: Argentina: Chubut (Ohaus 1909a).
Myloxena patagonica Ohaus, 1909
Original combination: Myloxena patagonica Ohaus, 1909: 440 (Ohaus 1909a).
Type locality: "Patagonien."
Distribution: Argentina: "Patagonia" (Ohaus 1909a).
Myloxena vestita Burmeister, 1881
Original combination: Myloxena vestita Burmeister, 1881: plate 2 (in Berg 1881).
Type locality: "la Pampa occidental" and "Sud-Oeste del Rio Colorado."
Distribution: Argentina: Río Negro (Berg 1881).
Genus Neogutierrezia Martínez, 1953
Neogutierrezia Martínez, 1953: 3 (Martínez 1953a).
Type species: Neogutierrezia mirablis Martínez, 1953 by original designation.
Neogutierrezia araucana Martínez, 1973
Original combination: Neoguttierrezia araucana Martínez, 1973: 36.
Type locality: "Argentina, provincia de Neuquén, Arroyo Picún Leufú y ruta nacional no 40."
Distribution: Argentina: Neuquén (Martínez 1973).
Neogutierrezia mirabilis affinis Martínez, 1973
Original combination: Neoguttierrezia mirabilis affinis Martínez, 1973: 35.
Type locality: "Argentina, provincia de Río Negro, San Antonio Oeste (Playa "Las Grutas")."
Distribution: Argentina: Río Negro (Martínez 1973).
Neogutierrezia mirabilis mirabilis Martínez, 1953
Original combination: Neoguttierrezia mirablis Martínez, 1953: 3 (Martínez 1953a).
Type locality: "Argentina, Territorio Nacional de Rio Negro, Coronel Gomez."
Distribution: Argentina: Río Negro (Martínez 1973).
Ptyophis Redtenbacher, 1868: 69.
Type species: Ptyophis macrophylla Redtenbacher, by monotypy.
Preoccupied synonym: Tetraphyllus Philippi, 1864: 448.
Type species: Tetraphyllus paulseni Philippi, by monotypy.
Ptyophis paulseni (Philippi, 1864)
Original combination: Tetraphyllus paulseni Philippi, 1864: 448.
Type locality: "cordillera de Santiago."
Distribution: Chile: Región Metropolitana (Philippi 1864).
Synonym: Ptyophis macrophylla Redtenbacher, 1868: 70.
Type locality: "Zwei Männchen aus Chili."
Type genus: Macrodactylus Dejean, 1821: 58.
Genus Astaenosiagum Martínez, 1957
Astaenosiagum Martínez, 1957: 50.
Type species: Schizochelus longipes Philippi, 1861 by original designation.
Astaenosiagum longipes (Philippi, 1861)
Original combination: Schizochelus longipes Philippi, 1861: 738.
Type locality: "Valdivia."
Distribution: Chile: X Región de Los Lagos.
Genus Chremastodus Solier, 1851
Chremastodus Solier, 1851: 103.
Type species: Chremastodus pubescens Solier, 1851 by subsequent designation (Evans 2003).
Remark: misspelled as Cremastodus Solier by Dalla Torre (1913: 328).
Chremastodus marmoratus (Curtis, 1844)
Original combination: Macrodactylus marmoratus Curtis, 1844: 200.
Type locality: "Chiloe."
Chremastodus pubescens Solier, 1851
Original combination: Chremastodus pubescens Solier, 1851: 103.
Type locality: "la provincia de Coquimbo."
Genus Issacaris Fairmaire, 1889
Issacaris Fairmaire, 1889: cxvii.
Type species: Issacaris petalophora Fairmaire (a synonym of Dicrania aeneobrunnea Philippi), by monotypy.
Issacaris bullocki Gutiérrez, 1952
Original combination: Issacaris bullocki Gutiérrez, 1952: 221.
Type locality: "Chile: Prov. de Malleco: Angol; Cerros de Nahuelbuta, Angol."
Distribution: Chile: IX Región de La Araucania (Gutiérrez 1952).
Issacaris petalophora Fairmaire, 1889
Original combination: Issacaris petalophora Fairmaire, 1889: cxviii.
Type locality: "Chili."
Distribution: Chile: X Región de Los Lagos (Gutiérrez 1946a).
Issacaris setosiventris Gutiérrez, 1952
Original combination: Issacaris setosiventris Gutiérrez, 1952: 219.
Type locality: "Chile: Prov. Osorno: Puyehue."
Distribution: Chile: X Región de Los Lagos (Gutiérrez 1952).
Genus Macrodactylus Dejean, 1821
Macrodactylus Dejean, 1821: 58.
Type species: Melolontha subspinosa Fabricius, by subsequent designation (Evans 2003).
Synonym: Stenothorax Harris, 1827: 8.
Type species: Melolontha subspinosa Fabricius, by monotypy.
Macrodactylus chilensis Solier, 1851
Original combination: Macrodactylus chilensis Solier, 1851: 115.
Type locality: "la provincia de Coquimbo."
Distribution: Argentina: Neuquén (Roig-Juñent et al. 2005); Chile: IV Región de Coquimbo (Gutiérrez 1949).
Macrodactylus crassipes Philippi, 1864
Original combination: Macrodactylus crassipes Philippi, 1864: 444.
Type locality: "Chile."
Distribution: Chile: VIII Región del Biobío (Gutiérrez 1949).
Synonym: Macrodactylus nigrinus Philippi, 1864: 443.
Type locality: "Chile."
Macrodactylus farinosus Philippi, 1864
Original combination: Macrodactylus farinosus Philippi, 1864: 442.
Type locality: "Andes prov. de Lineares."
Distribution: Chile: VII Región del Maule (Philippi 1864).
Genus Modialis Fairmaire and Germain. 1860
Modialis Fairmaire and Germain, 1860: 267.
Type species: Modialis prasinella Fairmaire and Germain, by monotypy.
Synonym: Acanthosternum Philippi, 1861: 739.
Type species: Acanthosternum splendens Philippi, by monotypy.
Modialis prasinella Fairmaire and Germain, 1860
Original combination: Modialis prasinella Fairmaire and Germain, 1860: 267.
Type locality: "Valdivia."
Distribution: Argentina: Neuquén; Chile: VIII Región del Biobío, X Región de Los Lagos (Gutiérrez 1947, 1949; Fairmaire and Germain 1860).
Synonym: Acanthosternum splendens Philippi, 1861: 739.
Type locality: "Valdivia."
Genus Phytholaema Blanchard, 1851
Phytholaema Blanchard, 1851: 218.
Type species: Areoda mutabilis Solier, 1851 by monotypy.
Remarks: Areoda Solier, 1851: 92 (not MacLeay, 1819) is a misapplication. Key to species in Saylor, 1937a: 8.
Synonym: Lacris Fairmaire and Germain, 1860: 268.
Type species: Lacris dilutipes Fairmaire and Germain, 1860 by monotypy.
Synonym: Melicurus Germain, 1905: 470.
Type species: Phytholaema herrmanni Germain, 1901 by subsequent designation (Smith 2002).
Phytholaema dilutipes (Fairmaire and Germain, 1860)
Original combination: Lacris dilutipes Fairmaire and Germain, 1860: 268.
Type locality: "Chillan."
Distribution: Chile: VII Región del Maule, VIII Región del Biobío (Gutiérrez 1944).
Synonym: Phytholaema flavipes Philippi, 1861: 735.
Type locality: "la cordillera de Linares."
Synonym: Phytholaema elaphocera Redtenbacher, 1868: 65.
Type locality: "Chili."
Phytholaema herrmanni herrmanni Germain, 1901
Original combination: Phytholaema herrmanni Germain, 1901: 10.
Type locality: "Araucanía setentrional."
Distribution: Chile: VII Región del Maule (Gutiérrez 1944).
Phytholaema herrmanni pallida Saylor, 1937
Original combination: Phytholaema herrmanni pallida Saylor, 1937a: 10.
Type locality: "Southern Chile."
Distribution: Chile (Saylor 1937a).
Synonym: Phytholaema peccans Blackwelder, 1944: 221.
Type locality: "Southern Chile."
Remark: Blackwelder (1944) erected this replacement name for Phytholaema pallida Porter, 1939. However, Blackwelder was erroneous when he determined that the Porter name was available.
Phytholaema mutabilis (Solier, 1851)
Original combination: Areoda mutabilis Solier, 1851: 93.
Type locality: "Valdivia."
Distribution: Argentina: Río Negro; Chile: VII-X Región (Gutiérrez 1944).
Plectris Le Peletier and Audinet-Serville, 1828: 369.
Type species: Plectris tomentosa Le Peletier and Audinet-Serville, by monotypy.
Synonym: Pseudoserica Guérin-Méneville, 1838: 86.
Type species: Pseudoserica marmorea Guérin-Méneville, by monotypy.
Synonym: Euryaspis Blanchard, 1851: 130.
Type species: Euryaspis gaudichaudii Blanchard, by monotypy.
Synonym: Trichoderma Nonfried, 1894: 11.
Type species: Trichoderma ceylanica Nonfried, by monotypy.
Synonym: Junkia Dalla Torre, 1913: 310.
Type species: Trichoderma ceylanica Nonfried, by monotypy.
Plectris blanchardi Frey, 1967
Original combination: Plectris blanchardi Frey, 1967a: 86 (replacement name for Plectris argentina Moser 1926).
Type locality: "Argentina: San Ignacio."
Distribution: Argentina: Neuquén (Moser 1926).
Synonym: Plectris argentina Moser 1926: 206 (secondary junior homonym of Plectris argentina [Bruch, 1909]).
Type locality: "Argentina: San Ignacio."
Genus Pristerophora Harold, 1869
Pristerophora Harold, 1869: 123 (Harold 1869c) (replacement name for Prionophora Solier, 1851: 101).
Type species: Prionophora picipennis Solier, by monotypy.
Preoccupied synonym: Prionophora Solier, 1851: 101.
Type species: Prionophora picipennis Solier, by monotypy.
Pristerophora flavipennis (Philippi, 1864)
Original combination: Prionophora flavipennis Philippi, 1864: 436.
Type locality: "E provintia Valdivia?"
Distribution: Chile: X Región de Los Lagos (Philippi 1864).
Pristerophora picipennis (Solier, 1851)
Original combination: Prionophora picipennis Solier, 1851: 102.
Type locality: "la provincia de Coquimbo."
Distribution: Chile: IV Región de Coquimbo (Solier 1851), VIII Región del Biobío (Gutiérrez 1949).
Pseudodicrania Gutiérrez, 1950a: 275.
Type species: Dicrania aeneobrunnea Philippi, by original designation.
Pseudodicrania aeneobrunnea (Philippi, 1861)
Original combination: Dicrania aeneobrunnea Philippi, 1861: 738.
Type locality: "Valdivia."
Distribution: Chile: X Región de Los Lagos (Philippi 1861).
Genus Schizochelus Blanchard, 1850
Schizochelus Blanchard, 1850: 89.
Type species: Schizochelus flavescens Blanchard, by subsequent designation (Evans 2003).
Schizochelus breviventris Philippi, 1864
Original combination: Schizochelus breviventris Philippi, 1864: 445.
Type locality: "Chile."
Distribution: Chile (Philippi 1864).
Schizochelus modestus Philippi, 1861
Original combination: Schizochelus modestus Philippi, 1861: 738.
Type locality: "Valdivia."
Distribution: Chile: X Región de Los Lagos (Philippi 1861).
Schizochelus serratus Philippi, 1864
Original combination: Schizochelus serratus Philippi, 1864: 444.
Type locality: "Chile."
Distribution: Argentina: Río Negro; Chile: X Región de Los Lagos (Gutiérrez 1947).
Schizochelus ursulus Philippi, 1864
Original combination: Schizochelus ursulus Philippi, 1864: 446.
Type locality: "Valdivia."
Distribution: Chile: X Región de Los Lagos (Philippi 1864).
Schizochelus vestitus Philippi, 1864
Original combination: Schizochelus vestitus Philippi, 1864: 447.
Type locality: "Valdivia."
Distribution: Chile: X Región de Los Lagos (Philippi 1864).
Type genus: Rutela Latreille, 1802: 151.
Type genus: Rutela Latreille, 1802: 151.
Genus Oogenius (Oogenius) Solier, 1851
Oogenius Solier, 1851: 97.
Type species: Oogenius virens Solier, by monotypy.
Remark: Although Oogeneius is the original spelling of this generic name, the incorrect subsequent spelling of Oogenius has come into prevailing usage. Therefore Oogenius should be considered to be the valid spelling.
Oogenius (Oogenius) arrowi Gutiérrez, 1949
Original combination: Oogenius arrowi Gutiérrez, 1949: 27.
Type locality: "Argentina... Puenta del Inca (Mendoza)."
Distribution: Argentina: Mendoza (Gutiérrez 1949).
Oogenius (Oogenius) castilloi Martínez and Peña, 1990
Original combination: Oogenius castilloi Martínez and Peña, 1990: 9.
Type locality: "Chile, provincia Ovalle, Fray Jorge."
Distribution: IV Región de Coquimbo (Martínez and Peña 1990).
Oogenius (Oogenius) chilensis barrosi Gutiérrez, 1949
Synonym: Oogenius barrosi Gutiérrez, 1949: 27.
Type locality: "Provincia de Santiago, Alhue (Cordillera de la Costa)."
Distribution: Región Metropolitana (Gutiérrez 1949).
Oogenius (Oogenius) chilensis chilensis Ohaus, 1905
Original combination: Oogenius chilensis Ohaus, 1905: 326 (Ohaus 1905b).
Type locality: "Chile, Valparaiso, Valdivia."
Distribution: Chile: VI-VIII Región (Gutiérrez 1949).
Oogenius (Oogenius) kuscheli Gutiérrez, 1949
Original combination: Oogenius kuscheli Gutiérrez, 1949: 29.
Type locality: "Volcán Copahue, Provincia de Bío-Bío, Chile."
Distribution: Chile: VIII Región del Biobío (Gutiérrez 1949).
Oogenius (Oogenius) lariosae Martínez, 1954
Original combination: Oogenius lariosae Martínez, 1954: 77 (Martínez 1954a).
Type locality: "Argentina, Provincia de Mendoza, Cordillera de los Andes, Paralelo."
Distribution: Argentina: Mendoza (Martínez 1954a).
Oogenius (Oogenius) penai Mondaca, 2005
Original combination: Oogenius penai Mondaca, 2005: 19.
Type locality: "Laderas sur orientales del cerro de El Roble, Caleu, Chacabuco" (Chile).
Distribution: Chile.
Remark: Replacement name for Oogenius gutierrezi Martínez and Peña, 1994 (a primary junior homonym of Oogenius gutierrezi Martínez, 1954).
Synonym: Oogenius gutierrezi Martínez and Peña, 1994: 117.
Type locality: "Laderas sur orientales del cerro de El Roble, Caleu, Chacabuco" (Chile).
Oogenius (Oogenius) virens Solier, 1851
Original combination: Oogenius virens Solier, 1851: 98.
Type locality: "escasamente en Chile."
Distribution: Chile: IV Región de Coquimbo, V Región de Valparaíso (Mondaca 2005).
Genus Oryctomorphus Guérin-Méneville, 1831
Oryctomorphus Guérin-Méneville, 1831b: plate 3.
Type species: Oryctomorphus bimaculatus Guérin-Méneville, 1831 by monotypy.
Synonym: Gonocheile Waterhouse, 1842: 281.
Type species: Oryctomorphus pictus Waterhouse, 1842 by monotypy.
Oryctomorphus bimaculatus Guérin-Méneville, 1831
Original combination: Oryctomorphus bimaculatus Guérin-Méneville, 1831b: plate 3.
Type locality: "Conception, au Chili."
Distribution: Argentina: Neuquén; Chile: IV Región de Coquimbo, V Región de Valparaíso, VIII Región del Biobío, X Región de Los Lagos (Gutiérrez 1947, 1949; Endrödi 1969).
Synonym: Oryctomorphus variegatus Guérin-Méneville, 1838: 52.
Type locality: "Pérou."
Synonym: Oryctomorphus pictus Waterhouse, 1842: 281.
Type locality: "Valdivia."
Oryctomorphus laevipennis Germain, 1855
Original combination: Oryctomorphus laevipennis Germain, 1855: 397.
Type locality: not given (Chile).
Distribution: Chile: V Región de Valparaíso, VIII Región del Biobío (Gutiérrez 1949, Endrödi 1969).
Synonym: Oryctomorphus parumstriatus Fairmaire, 1856: 483.
Type locality: "Chilensia."
Synonym: Oryctomorphus fairmairei Redtenbacher, 1868: 76.
Type locality: "Chili."
Oryctomorphus maculicollis Guérin-Méneville, 1838
Original combination: Oryctomorphus maculicollis Guérin-Méneville, 1838: 53.
Type locality: "le Pérou, près de Lima" (erroneous).
Distribution: Chile: VII Región del Maule (Endrödi 1969).
Remark: The record from Perú by Ohaus (1952) is dubious and should be disregarded.
Synonym: Oryctomorphus morio Burmeister, 1847: 31.
Type locality: "Chili."
Type genus: Anoplognathus Leach, 1815: 156.
Type genus: Brachysternus Guérin-Méneville, 1831: plate 3.
Aulacopalpus Guérin-Méneville, 1838: 57.
Type species: Aulacopalpus viridis Guérin-Méneville, by monotypy.
Remark: key to species in Smith 2002.
Synonym: Tribostethes Curtis, 1844: 197.
Type species: Brachysternus castaneus Laporte, by monotypy.
Synonym: Bembegeneius Solier 1851: 84.
Type species: Bembegeneius fulvescens Solier, by monotypy.
Synonym: Sulcipalpus Harold 1869: 1232 (Harold 1869a).
Type species: Aulacopalpus viridis Guérin-Méneville, by monotypy.
Synonym: Pseudadelphus Arrow 1899: 369.
Type species: Tribostethes ciliatus Solier, by monotypy.
Synonym: Amblyterodes Germain 1905: 470.
Type species: Aulacopalpus viridis Guérin-Méneville, by monotypy.
Synonym: Aegolasia Kolbe 1907: 112.
Type species: Aegolasia michaelseni Kolbe, by monotypy.
Aulacopalpus aconcaguensis Smith, 2002
Original combination: Aulacopalpus aconcaguensis Smith, 2002: 401.
Type locality: "Chile: Aconcagua Cachagua."
Distribution: Chile: IV Región de Coquimbo, V Región de Valparaíso, Región Metropolitana (Smith 2002).
Aulacopalpus castaneus (Laporte, 1840)
Original combination: Brachysternus castaneus Laporte, 1840: 127.
Type locality: "Chile Santiago."
Distribution: Chile: IV-IX Región, Región Metropolitana (Smith 2002).
Synonym: Bembegeneius fulvescens Solier 1851: 85.
Type locality: "Santiago, Concepcion y la Araucania."
Synonym: Tribostethes cupreus Philippi and Philippi 1864: 321.
Type locality: "Chilenischen."
Synonym: Aulacopalpus fulvovirens Ohaus 1909: 12 (Ohaus 1909b).
Type locality: "Chile."
Aulacopalpus ciliatus (Solier, 1851)
Original combination: Tribostethes ciliatus Solier 1851: 89.
Type locality: "Santiago, Concepcion y la Araucania."
Distribution: Chile: IV-IX Región, Región Metropolitana (Smith 2002).
Aulacopalpus clypealis Ohaus, 1905
Original combination: Aulacopalpus clypealis Ohaus 1905: 137 (Ohaus 1905a).
Type locality: "Chile."
Distribution: Chile: V Región de Valparaíso, VI Región del O'Higgins, Región Metropolitana (Smith 2002).
Aulacopalpus pilicollis (Fairmaire, 1884)
Original combination: Tribostethes pilicollis Fairmaire 1884: 491.
Type locality: "Magellan et de Santa-Cruz."
Distribution: Argentina: La Rioja, Neuquén, Río Negro, Chubut, Santa Cruz, Tierra del Fuego; Chile: IX Región de la Araucania, XI Región Aisen, XII Región de Magallanes (Smith 2002).
Synonym: Aegolasia michaelseni Kolbe 1907: 112.
Type locality: "Chile Magallanes" (based on neotype designated by Smith [2002]).
Aulacopalpus punctatus (Fairmaire and Germain, 1860)
Original combination: Tribostethes punctatus Fairmaire and Germain 1860: 268.
Type locality: "Chile Valdivia."
Distribution: Chile: X Región de Los Lagos (Smith 2002).
Synonym: Amblyterus variabilis Philippi 1861: 742.
Type locality: "Valdivia."
Aulacopalpus pygidialis Ohaus, 1905
Original combination: Aulacopalpus pygidialis Ohaus 1905:139 (Ohaus 1905a).
Type locality: "Chile."
Distribution: Chile: VIII Región del Biobío, X Región de Los Lagos (Smith 2002).
Aulacopalpus valdiviensis Smith, 2002
Original combination: Aulacopalpus valdiviensis Smith, 2002: 417.
Type locality: "Sn. Juan Corral Valdivia."
Distribution: Chile: X Región de Los Lagos (Smith 2002).
Aulacopalpus viridis Guérin-Méneville, 1838
Original combination: Aulacopalpus viridis Guérin-Méneville 1838a: 58.
Type locality: "Chile."
Distribution: Chile: VII-IX Región, Región Metropolitana (Smith 2002).
Synonym: Tribostethes virens Philippi and Philippi 1864: 320.
Type locality: "Tregualemu N. Costa Nuble."
Brachysternus Guérin-Méneville, 1831: plate 3.
Type species: Brachysternus prasinus Guérin-Méneville, by monotypy.
Remark: key to species in Jameson and Smith 2002.
Brachysternus angustus (Philippi and Philippi, 1864)
Original combination: Aulacopalpus angustus Philippi and Philippi, 1864: 322.
Type locality: "Chilenischen."
Distribution: Argentina: Neuquén; Chile: VI-X Región (Jameson and Smith 2002).
Brachysternus germaini (Ohaus, 1909)
Original combination: Tribostethes germaini Ohaus, 1909: 14 (Ohaus 1909b).
Type locality: "Chile Talca La Vega (E Carrizal)" (neotype designated by Jameson and Smith [2002]).
Distribution: Chile: VII Región del Maule, VIII Región del Biobío (Jameson and Smith 2002).
Brachysternus marginatus Germain, 1905
Original combination: Brachysternus marginatus Germain, 1905: 485.
Type locality: "Chile."
Distribution: Argentina: Neuquén; Chile: VII-IX Región (Jameson and Smith 2002, Roig-Juñent et al. 2005).
Brachysternus olivaceus Philippi and Philippi, 1864
Original combination: Brachysternus olivaceus Philippi and Philippi, 1864: 318.
Type locality: "Chilenischen."
Distribution: Chile: IV-VII, Región Metropolitana (Jameson and Smith 2002).
Synonym: Brachysternus chloris Philippi and Philippi, 1864: 319.
Type locality: "S. Fernando" (Chile).
Synonym: Brachysternus riverae Germain, 1905: 497.
Type locality: "proviene de los alrededores de Chillan."
Synonym: Brachysternus herbaceus Germain, 1905: 499.
Type locality: "Chile."
Synonym: Brachysternus araucanicus Ohaus, 1905: 156.
Type locality: "Araucania."
Brachysternus patagoniensis Jameson and Smith, 2002
Original combination: Brachysternus patagoniensis Jameson and Smith, 2002: 348.
Type locality: "Chile: 45 km W Angol Nahuelbuta Nat. Pk."
Distribution: Argentina: Neuquén, Chubut; Chile: IX-XI Región (Jameson and Smith 2002).
Brachysternus prasinus Guérin-Méneville, 1831
Original combination: Brachysternus prasinus Guérin-Méneville, 1831: plate 3.
Type locality: "Chile Valdivia."
Distribution: Argentina: Neuquén, Río Negro, Chubut; Chile: IV-X Región, Región Metropolitana (Jameson and Smith 2002).
Synonym: Brachysternus fulvipes Guérin-Méneville, 1838: 61.
Type locality: "Pérou."
Synonym: Brachysternus vicinus Guérin-Méneville, 1839: 300.
Type locality: "le détroit de Magellan."
Synonym: Brachysternus sinuatifrons Germain, 1905: 475.
Type locality: "proviene de los valles andinos de donde sale el rio Cachapoal."
Synonym: Brachysternus viridis Germain, 1905: 487.
Type locality: "Chile."
Synonym: Brachysternus pubescens Germain, 1905: 495.
Type locality: "Chile."
Synonym: Brachysternus dilatatus Germain, 1905: 502.
Type locality: "Chile."
Synonym: Brachysternus viridipes Ohaus, 1905: 157.
Type locality: "Chile Victoria" (based on lectotype) (Jameson and Smith 2002).
Synonym: Brachysternus hirtus Ohaus, 1905: 158.
Type locality: "Osorno S. Chile" (based on lectotype) (Jameson and Smith 2002).
Brachysternus spectabilis Erichson, 1847
Original combination: Brachysternus spectabilis Erichson, 1847: 100 (Erichson 1847a).
Type locality: "Republica Peruana" (erroneous) emended to "Chile or Argentina" by Jameson and Smith (2002).
Distribution: Argentina: Neuquén, Río Negro, Chubut; Chile: V-XI Región, Región Metropolitana (Jameson and Smith 2002).
Synonym: Brachysternus major Philippi and Philippi, 1864: 317.
Type locality: "Chilenischen."
Synonym: Brachysternus obscurus Philippi and Philippi, 1864: 317.
Type locality: "Chilenischen."
Synonym: Brachysternus philippii Germain, 1905: 481.
Type locality: "Chile."
Hylamorpha Arrow, 1899: 368.
Type species: Aulacopalpus elegans Burmeister, 1844 by subsequent designation (Ratcliffe and Ocampo 2002).
Hylamorpha elegans (Burmeister, 1844)
Original combination: Aulacopalpus elegans Burmeister, 1844: 459.
Type locality: "Santiago, Chile."
Distribution: Argentina: Neuquén, Río Negro, Chubut; Chile: IV Región de Coquimbo, V Región de Valparaíso, VII-X Región, Región Metropolitana (Ratcliffe and Ocampo 2002).
Synonym: Callichloris perelegans Curtis, 1844: 198.
Type locality: "Valparaiso."
Synonym: Sulcipalpus subviolaceus Nonfried, 1894: 125.
Type locality: "Cordilleras de Chile."
Synonym: Hylamorpha cylindrica Arrow, 1899: 368.
Type locality: "Chili."
Synonym: Hylamorpha rufimana Arrow, 1899: 368.
Type locality: "Chili."
Type genus: Dynastes MacLeay, 1819: 22.
Type genus: Cyclocephala Dejean, 1821: 57.
Genus Cyclocephala Dejean, 1821
Cyclocephala Dejean, 1821: 57.
Type species: not verified.
Synonym: Aclinidia Casey, 1915: 113.
Type species: Melolontha castanea Fabricius by original designation.
Synonym: Diapatalia Casey, 1915: 111.
Type species: Cyclocephala discicollis Arrow by original designation.
Synonym: Dichromina Casey, 1915: 112.
Type species: Cyclocephala dimidiata Burmeister by original designation.
Synonym: Graphalia Casey, 1915: 159.
Type species: not verified.
Synonym: Halotosia Casey, 1915: 113.
Type species: Cyclocephala fasciolata Bates, 1888 by original designation.
Synonym: Homochromina Casey, 1915: 113.
Type species: Homochromina divisa Casey, 1915 by original designation.
Synonym: Isocoryna Casey, 1915: 136.
Type species: Cyclocephala jalapensis Casey, 1915 by monotypy.
Synonym: Mononidia Casey, 1915: 110.
Type species: Cyclocephala carbonaria Arrow by original designation.
Synonym: Ochrosidia Casey, 1915: 112.
Type species: Melolontha immaculata Olivier, 1789 by original designation.
Synonym: Plagiosalia Casey, 1915: 135.
Type species: not verified.
Synonym: Spilosota Casey, 1915: 112.
Type species: Spilosota nubecula Casey, 1915 by original designation.
Synonym: Stigmalia Casey, 1915: 111.
Type species: Cyclocephala mafaffa Burmeister, 1847 by original designation.
Synonym: Aspidolites Höhne, 1922: 374.
Type species: Aspidolea atricollis Höhne, 1922 by monotypy.
Synonym: Aspidolella Prell, 1936: 146 (replacement name for Aspidolites Höhne, 1922).
Type species: Aspidolea atricollis Höhne, 1922 by monotypy.
Synonym: Albridarollia Bolívar y Pieltain, Jiménez-Asúa, and Martínez, 1963: 182.
Type species: Albridarollia ocellata Bolívar y Pieltain, Jiménez-Asúa, and Martínez, 1963 by original designation.
Synonym: Paraclinidia Martínez, 1965.
Type species: not verified.
Cyclocephala metrica Steinheil, 1874
Original combination: Cyclocephala metrica Steinheil, 1874: 559.
Type locality: "Rio Cuarto e San Luis."
Distribution: Uruguay; Argentina: Tucumán, La Pampa, San Luis, Buenos Aires, Río Negro (Bruch 1911, Endrödi 1966).
Remark: The country record for México in Endrödi (1966) is dubious and should be disregarded.
Synonym: Cyclocephala parvula Burmeister, 1881: plate 2 (in Berg 1881).
Type locality: "Salinas Chicas" and "Banda Oriental."
Genus Dyscinetus Harold, 1869
Dyscinetus Harold, 1869: 123 (replacement name for Chalepus MacLeay, 1819).
Type species: Melolontha geminata Fabricius, 1801 by monotypy.
Synonym: Chalepus MacLeay, 1819: 149.
Type species: Melolontha geminata Fabricius, 1801 by monotypy.
Synonym: Palechus Casey, 1915: 174.
Type species: Dyscinetus histrio Casey, 1915 by original designation.
Dyscinetus australis Joly and Escalona, 2002
Original combination: Dyscinetus australis Joly and Escalona, 2002: 202.
Type locality: "Argentina: Neuquen: Convuco, Usina."
Distribution: Argentina: Buenos Aires, Neuquén, Río Negro (Joly and Escalona 2002).
Type genus: Pentodon Hope, 1837: 30.
Genus Bothynus Hope, 1837
Bothynus Hope, 1837: 30.
Type species: Geotrupes cuniculus Fabricius, 1801 by original designation.
Synonym: Cratocnemis Blanchard, 1842: plate 11.
Type species: Cratocnemis niger Blanchard, 1842 by monotypy.
Synonym: Scaptophilus Burmeister, 1847: 122.
Type species: not verified.
Synonym: Corynoscelis Burmeister, 1847: 126.
Type species: not verified.
Bothynus minor Steinheil, 1874
Original combination: Bothynus minor Steinheil, 1874: 561.
Type locality: "Patagones."
Distribution: Argentina: Chaco, "Patagones" (Bruch 1911).
Genus Thronistes Burmeister, 1847
Thronistes Burmeister, 1847: 177.
Type species: Thronistes rouxii Burmeister, 1847 by monotypy.
Thronistes niger Blanchard, 1842
Original combination: Cratocnemis niger Blanchard, 1842: plate 11.
Type locality: "les dunes de la baie de San-Blas (Patagonie)."
Distribution: Argentina: Chubut (Blanchard 1847).
Genus Tomarus Erichson, 1847
Tomarus Erichson, 1847: 95 (Erichson 1847a).
Type species: Tomarus maimon Erichson, 1847 by monotypy.
Tomarus villosus Burmeister, 1847
Original combination: Podalgus villosus Burmeister, 1847: 120.
Type locality: "Chili und dem westlichen Patagonien."
Distribution: Peru; Uruguay; Argentina: Buenos Aires, Mendoza, Río Negro; Chile: Region IV de Coquimbo, V Región de Valparaíso, Región Metropolitana, VIII Región del Biobío, X Región de Los Lagos (Bruch 1911; Gutiérrez 1945, 1947; Endrödi 1969).
Synonym: Oryctes nitidicollis Solier 1851: 79.
Type locality: "Santa Rosa, Santiago y Coquimbo."
Synonym: Ligyrus patagonus Steinheil, 1874: 560.
Type locality: "Patagones."
References cited
Allsopp, P. G. 1983. Neophaenognatha, a new genus for the Neotropical species of Phaenognatha Hope (Scarabaeidae: Aclopinae) with the description of N. capella n. sp. and designation of lectotypes. Coloepterists Bulletin 37(3): 208-211.
Arrow, G. J. 1899. On the classification of the coleopterous family Rutelidae. Annals and Magazine of Natural History (4)7: 363-370.
Arrow, G. J. 1901. Remarks on the secondary sexual differences in rutelid Coleoptera, with descriptions of some new forms. Annals and Magazine of Natural History (7)7: 393-401.
Arrow, G. J. 1903. On the Laparostict Lamellicorn Coleoptera of Grenada and St. Vincent. Transaction of the Entomological Society of London 1903: 509-520.
Arrow, G. J. 1904. Sound-production in the Lamellicorn beetles. Transactions of the Entomological Society of London 1904: 709-750.
Arrow, G. J. 1940. A nomenclatural note (Coleoptera). Proceedings of the Royal Entomological Society of London. Series B. Taxonomy 9: 16.
Audouin, M. V. and M. A. Brullé. 1837. Histoire Naturelle des Insectes, volume 6. Coléoptères 3. F. D. Pillot, Paris. 448 pp.
Balthasar, V. 1939. Eine Vorstudie zur Monographie der Gattung Canthon Hffsg. (10. Beitrag zur Kenntnis der Scarabaeiden der neotropischen Region). Folia Zoologica et Hydrobiologica 9: 179-238.
Balthasar, V. 1939. Neue Arten der coprophagen Scarabaeiden aus dem Museo Zoologico delle R.Università di Firenze. Redia 25: 1-36.
Balthasar, V. 1941. Gli Scarabeidi coprofagi e Trogini dell'Africa orient. Ital. del Museo di Milano. Atti de la Societa Italiana di Scienze Naturali del Museo Civico di Soria 80: 103-154.
Balthasar, V. 1942. Neue Arten und Gattungen der coprophagen Scarabaeiden. 75. Beitrag zur Kenntnis der Scarabaeiden (Col.). Acta Entomologica Musaei Pragae 20: 188-206.
Balthasar, V. 1946. De novis generis Aphodius Illig. speciebus. Acta Entomologica Musei Nationalis Pragae 24: 53-68.
Balthasar, V. 1947. De novis generis Ataenius Har. speciebus. Acta Entomologica Musei Nationalis Pragae 25: 49-54.
Balthasar, V. 1959. Beitrag zur kenntniss des gattung Onthophagus Latr. Acta entomologica Musei Nationalis Pragae 33(565): 461-471.
Balthasar, V. 1960. Beiträge zur Kenntnis der Insektenfauna Boliviens Teil XIII. Coleoptera XII. Neue Aphodiinae-Arten (100. Beitrag zur Kenntnis der Scarabaeidae). Opuscula Zoologica 48: 1-7.
Balthasar, V. 1961. Eine neue Gattung und neue Arten der Unterfamilie Aphodiinae. Deutsche Entomologische Zeitschrift, Neue Folge 8(1/2): 121-130.
Balthasar, V. 1963. Eine neue Gattung, Untergattung und neue Arten der Familie Aphodiidae (Coleoptera). Reichenbachia 1: 277-290.
Bates, H. W. 1887. Pectinicornia and Lamellicornia, pp 25-160. In F. D. Godman and O. Salvin (eds.), Biologia Centrali-Americana. Insecta. Coleoptera, volume 2, part 2. 432 pp, 24 plates.
Bates, H. W. 1888. Pectinicornia and Lamellicornia, pp 161-342. In F. D. Godman and O. Salvin (eds.), Biologia Centrali-Americana. Insecta. Coleoptera, volume 2, part 2. 432 pp, 24 plates.
Benesh, B. 1955. Some notes on Neotropical stagbeetles (Coleoptera: Lucanidae). Entomological News 46: 97-104.
Benesh, B. 1956. Some notes on Neotropical stagbeetles, with descriptions of new species (Coleoptera: Lucanidae). Entomological News 47: 57-68.
Berg, C. 1881. Insectos, pp 77-115. In: Informe Oficial de la Comision Cientifíca Agregada al Estado Mayor General de la Expedicion al Rio Negro (Patagonia). Realizada en los Meses de Abril, Mayo y Junio de 1879, Bajo las rdenes del General D. Julio A. Roca, entrega 1. Zoologica. Ostwald y Martinez, Buenos Aires.
Blackburn, T. 1892. Further notes on Australian Coleoptera, with descriptions of new genera and species. XI. Transactions of the Royal Society of South Australia 15: 20-73.
Blackburn, T. 1904. Further notes on Australian Coleoptera, with descriptions of new genera and species. XXXIV. Transactions of the Royal Society of South Australia 28: 281-297.
Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. United States National Museum Bulletin 185(2): 189-341.
Blanchard, C. É. 1841. Insectes de l'Amérique Méridionale. In A. d'Orbigny, C. É. Blanchard, and A. Brullé, Voyage dans l'Amérique Méridionale, volume 6, part 2. P. Bertrand, Paris. P. Bertrand, Paris. Plate 10.
Blanchard, C. É. 1842. Insectes de l'Amérique Méridionale. In A. d'Orbigny, C. É. Blanchard, and A. Brullé, Voyage dans l'Amérique Méridionale, volume 6, part 2. P. Bertrand, Paris. Plates 11-12.
Blanchard, C. É. 1845a. Histoire Naturelle des Insectes, Leurs Moeurs, Leurs Métamorphoses et Leur Classification ou Traité Élémentaire d'Entomologie, vol. 1. Librairie F. Savy, Paris. 398 pp.
Blanchard, C. É. 1845b. Insectes de l'Amérique Méridionale. In A. d'Orbigny, C. É. Blanchard, and A. Brullé, Voyage dans l'Amérique Méridionale, volume 6, part 2, livraison 78-80. P. Bertrand, Paris. Pp 113-184.
Blanchard, C. É. 1846. Insectes coléoptères. In J. B. Hombron and H. Jacquinot, Atlas d'histoire Naturelle Zoologie. Voyage au Pole Sud et dans l'Océanie sur les Corvettes l'Astrolabe et la Zélée; Exécuté par Ordre du Roi pendant les Années 1837-1838-1839-1840. Gide et J. Baudry, Paris. Plate 8.
Blanchard, C. É. 1847. Insectes de l'Amérique Méridionale. In A. d'Orbigny, C. É. Blanchard, and A. Brullé, Voyage dans l'Amérique Méridionale, volume 6, part 2, livraison 90. P. Bertrand, Paris. Pp 185-232.
Blanchard, C. E. 1850. Ordre des Coléoptéres. In: Milne-Edwards, H., Blanchard, E. and Lucas, H. (editors) Museum d'Histoire Naturelle de Paris. Catalogue de la collection entomologique. Classe des insectes. Gide and Baudry, Paris. pp. 1-128.
Blanchard, C. E. 1851. Ordre des Coléoptéres. In: Milne-Edwards, H., Blanchard, E. and Lucas, H. (editors) Museum d'Histoire Naturelle de Paris. Catalogue de la collection entomologique. Classe des insectes. Gide and Baudry, Paris. pp. 129-240.
Blanchard, C. É. 1854. Zoologie. Description des Insectes. In J. B. Hombron and H. Jacquinot, Voyage au Pole Sud et dans l'Océanie sur les Corvettes l'Astrolabe et la Zélée; Exécuté par Ordre du Roi pendant les Années 1837-1838-1839-1840. Zoologie, volume 4. Gide et J. Baudry, Paris. 422 pp.
Bolívar y Pieltain, C., L. Jiménez-Asúa, and A. Martínez. 1963. Notas sobre Dynastinae Neotropicales con especial referencia a especies Mexicanas (Col., Scarab.). Ciencia 22(6): 181-190.
Boucomont, A. 1911. Contribution a la classification de Geotrypidae [Col.]. Annales de la Société Entomologique de France 79: 333-350. [Dated 1910].
Brenske, E. 1900. Coléoptères. Scarabaeidae. In: Diagnoses d'insectes recuellis par l'expédition anatarctiques belge. Annales de la Société Entomologique de Belgique 44: 109-110.
Brenske, E. 1906. Coléoptères. Scarabaeidae. In: Expedition Antarctique Belge. Resultats du Voyage du S.Y. Belgica en 1897-1898-1899 sous le Commandment de A. de Gerlache de Gomery. Rapports Sientifiques. Publiés aux Frais du Government Belge, sous la Direction de la Commission de la Belgica. Zoologie. Insectes. J. E. Buschmann, Anvens. Pp. 29-34, plate 2.
Bruch, C. 1909. Nuevas especies de los géneros Philochloenia y Demodema (Coleópteros Lamelicornios). Revista del Museo de La Plata (2)16: 340-352, plate 2.
Bruch, C. 1910. Descripción de dos nuevos lamelicornios de la fauna Argentina. Revista del Museo de La Plata (2)17: 71-77.
Bruch, C. 1911. Catálogo Sistemático de los Coleópteros de la República Argentina. Pars IV. Familias Lucanidae, Scarabaeidae (Coprini-Cetonini), Passalidae. Revista del Museo de La Plata 17: 181-225.
Bruch, C. 1938. Algunas especies nuevas de coleópteros (Aphod. Lathrid. Nitidul.). Notas del Museo de La Plata. Zoología 3(12): 160-164.
Buquet, L. 1842. Description d'une espèce nouvelle de Lucanide, appartenant au genre Dorcus de Megerle. Annales de la Société Entomologique de France (2) 11: 283-284, plate 12.
Burmeister. H. C. C. 1844. Handbüch der Entomologie , volume 4, part 1. T.C.F. Enslin, Berlin, 586 pp.
Burmeister, H. 1846. III. Coleoptera. 7. Lamellicornia. 27. Copris, 4 unnumbered pages. In, H. Burmeister, Genera Quaedam Insectorum. Privately published, Berlin.
Burmeister. H. C. C. 1847. Handbüch der Entomologie, volume 5. T.C.F. Enslin, Berlin, 584 pp.
Burmeister, H. C. C. 1855. Handbuch der Entomologie, volume 4, part 2. T. C. F. Enslin, Berlin, 569 pp.
Burmeister, H. C. C. 1861. Die Ateuchiden ohne Fulskrallen, monographisch bearbeitet. Berliner Entomologische Zeitschrift 1861: 55-67.
Burmeister, H. C. C. 1873. Lamellicornia Argentina. Stettiner Entomologische Zeitung 34: 403-417.
Burmeister, H. C. C. 1874. Lamellicornia Argentina. Stettiner Entomologische Zeitung 35: 120-133.
Burmeister, H. C. C. 1876. Die Argentinischen Arten der Gattung Trox Fabr. Entomologische Zeitung 37: 241-268.
Burmeister, H. C. C. 1877. Die Argentinischen Aphodiaden. Entomologische Zeitung 38: 401-414.
Campos, L. and L. E. Peña. 1973. Los insectos de Isla de Pascua. Revista Chilena de Entomologia 7: 217-229.
Cartwright, O. L. 1946. Two new Psammodius from southeastern United States (Coleoptera: Scarabaeidae). Bulletin of the Brooklyn Entomological Society 41: 89-91.
Cartwright, O. L. 1955. Scarab beetles of the genus Psammodius in the Western Hemisphere. Proceedings of the United States National Museum 104(3344): 413-462.
Cekalovik, T. and M. Castro. 1983. Chiasognathus granti Stephens, 1831 (Coleoptera Lucanidae), descripción de la larva y nuevas localidades para la especie. Boletin de la Sociedad de Biologia de Concepción 54: 71-76.
Chalumeau, F. 1982. Contribution a l'etude des Scarabaeoidea des Antilles (III). Nouvelle Revue de Entomologique 12(4): 321-345.
Chalumeau, F. 1992. Eupariini du nouveau monde: un mise au point (Coleoptera, Scarabaeidae) (1re partie). Nouvelle Revue d'Entomologie 9: 189-206.
Chalumeau, F. and B. Brochier. 1995. Les Chiasognathinae: genres, sous-genres et synonymies (Coleoptera, Lucanidae). Bulletin de la Société Sciences Nat 83: 18-24.
Chalumeau, F. and B. Brochier. 2007. Chiasognathinae of the Andes. Taita Publishers, Hradec Králové, 324 pp.
Chapin, E. A. 1938. The nomenclature and taxonomy of the genera of the scarabaeid subfamily Glaphyrinae. Proceedings of the Biological Society of Washington 51: 79-86.
Chevrolat, L.A.A. 1844. In: Guérin-Méneville, F.E. Iconographie du régne animal de G. Cuvier... vol. 7, insectes 1829-1838. Paris. 576 pp.
Curtis, J. 1835. Characters and descriptions of a new genus of the family Melolonthidae. Transactions of the Zoological Society of London 1: 307-310.
Curtis, J. 1844. Descriptions of the insects collected by Capt. P. P. King, R. N., F. R. S., F. L. S. &c., in the survey of the Straits of Magellan. Proceedings of the Linnean Society of London 1(21): 196-202.
Curtis, J. 1845. Descriptions, &c. of the insects collected by Captain P. P. King, R. N., F. R. S. & L. S., in the survey of the Straits of Magellan. Transactions of the Linnean Society of London 19: 441-475.
von Dalla Torre, K.W. 1879. Systematisches Verzeichnis der in Oberösterreich bisher beobachteten Käfer. Jahres-Bericht des Vereines für Naturkunde in Oesterreich ob der Ens 10: 1-125.
von Dalla Torre, K.W. 1912. Coleopterorum catalogus, vol. 20, pars 47, Scarabaeidae; Melolonthidae II. pp. 85-134.
von Dalla Torre, K.W. 1913. Coleopterorum catalogus, vol. 20, pars 50, Scarabaeidae; Melolonthidae IV. pp. 291-450.
De Geer, C. 1774. Mémoires pour Servir à l'Histoire des Insects, volume 4. Pierre Hesselberg, Stockholm. 456 pp, 19 plates.
Dejean, P. F. M. A. 1821. Catalogue de la Collection de Coléoptères de M. le Baron Dejean. Crevot Libraire, Paris, 136 pp.
Dejean, P.F.M.A. 1833. Catalogue des Coléoptères de la collection de M. le Comte Dejean, Fascicles 1-2. Méquignon-Marvis Père et Fils, Paris, 176 pp.
Dellacasa, M. 1988. Contribution to a world-wide catalogue of Aegialiidae, Aphodiidae, Aulonocnemidae, Termitotrogidae (Coleoptera Scarabaeoidea). Memorie della Societa Entomologica Italiana 66: 1-455. [Dated 1987].
Dellacasa, G. 1992. On some little known species of Aphodius from Australia and South America with descriptions of a new subgenus. Frustula Entomologica 13(26): 145-158. [Dated 1990].
Dellacasa, M., R. D. Gordon, P. J. Harpootlian, Z. Stebnicka, and G. Dellacasa. 2002. Systematic redefinition of the New World Didactyliini (Coleoptera: Scarabaeidae: Aphodiinae) with descriptions of two new species of Aidophus Balthasar. Insecta Mundi 15(4): 193-216. [2001].
Dellacasa, M. and Z. Stebnicka. 2001. A new genus for Oxyomus morosus Harold, 1869 (Eupariini) and redefinition of Oxyomus setosopunctatus A. Schmidt, 1911 (Aphodiini) (Coleoptera Scarabaeidae: Aphodiinae). Acta Zoologica Mexicana (Nueva Serie) 83: 29-34.
Dellacasa, M. and Z. Stebnicka. 2001. Acanthaphodius bruchi A. Schmidt, 1909, femina nondum cognita (Coleoptera: Scarabaeidae: Aphodiinae). Acta Zoologica Cracoviensia 44(4): 431-433.
Dury, C. 1902. A revised list of the Coleoptera observed near Cincinnati, Ohio, with notes on localities, bibliographical references, and description of new species. Journal of the Cincinnati Society of Natural History 20: 107-189.
Endrödi, S. 1966. Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen Staatliches Museum für Tierkunde in Dresden 33:1-457.
Endrödi, S. 1969. Monographie der Dynastinae 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). Entomologische Abhandlungen Staatliches Museum für Tierkunde in Dresden 37: 1-145.
Endrödi, S. 1977. Monographie der Dynastinae 8. Tribus: Phileurini, amerikanische Arten I (Coleoptera). Folia Entomologica Hungarica 31: 7-45.
Erichson, W.F. 1835. Neue Südamerikanische Käfergattungen aus der familie der Blätterhörner. Archiv für Naturgeschichte 1: 256-270.
Erichson, W.F. 1847a. Conspectus insectorum coleopterorum quae in Republica Peruana observata sunt. Archiv für Naturgeschichte 13: 67-185.
Erichson, W. F. 1847b. Naturgeschichte der Insecten Deutschlands, volume 3, part 5. Nicolaischen Buchhandlung, Berlin, pp. 641-800.
Eschscholtz, J. F. 1822. Entomographien. O. Reimer, Berlin. 128 pp.
Evans, A. V. 2003. A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 211: 1-458.
Fabricius, J. C. 1775. Systema entomologiae sistens insectorum classes, ordines, genera, species, adjectis synonimis, locis, observationibus, diescriptionbus. Korte, Flensburg and Leipzig. 832 pp.
Fabricius, J. C. 1787. Mantissa Insectorum Sistens Eorum Species Nuper Detectas Adiectis Characteribus Genericis, Differentiis Specificis, Emendationibus, Observationibus, volume 1. Christ Gottl Proft, Copenhagen. 348 pp.
Fabricius, J. C. 1792. Entomologia Systematica Emendata et Aucta. Secundum Classes, Ordines, Genera, Species Adjectis Synonimis, Locis, Observationibus, Descriptionibus, volume 1. Christ Gottl Proft, Copenhagen. 330 pp.
Fabricius, J. C. 1798. Supplementum Entomologiae Systematicae. Proft et Storgh, Copenhagen. 572 pp.
Fabricius, J. C. 1801. Systema Eleutheratorum Secundum Ordines, Genera, Species Adiectis Synonymis, Locis, Observationibus, Descriptionibus, volume 1. Bibliopolii Academici Novi, Kiel. 506 pp.
Fairmaire, L. 1850. Description d'un nouveau genre de lucanide. Annales de la Société Entomologique de France series 2, 8: 53-57.
Fairmaire, L. 1856. Coleoptera Chilensia a Germain detecta et a Léon Fairmaire descripta. Revue et Magasin de Zoologie 2(8): 483-485.
Fairmaire, L. 1884. Note sur quelques Coléoptères de Magellan et de Santa Cruz. Annales de la Société Entomologique de France (6)3: 483-506.
Fairmaire, L. 1885. Description de trios nouvelles espèces de coléoptères. Bulletin de la Société Entomologique de France 1885: CLXXXIX.
Fairmaire, L. 1889. Coléoptères. In: Recherches sur les insectes recueilles pendant la mission chargée d'observer a Santa-Cruz de Patagonie le passage de Vénus. Nouvelles Archives du Muséum d'Histoire Naturelle (3)1: 101-137, plate 9.
Fairmaire, L. 1889. Les descriptions suivantes de nouvelles espèces de Coléoptères. Bulletin de la Société Entomologique de France 1889: cxvii-cxix.
Fairmaire, L. 1892. Descriptions de quelques Coléoptères Argentins. Annales de la Société Entomologique de Belgique 36: 242-253.
Fairmaire, L. 1893. Quelques coléoptères de la République Argentine. Annales de la Société Entomologique de Belgique 37: 608-612.
Fairmaire, L. and P. Germain. 1860. Coleoptera chilensia. Revue et Magasin de Zoologie (2)12: 267-269.
Fairmaire, L. and P. Germain. 1861. Coleoptera Chilensia, part 2. Félix Malteste, Paris. 8 pp.
Fairmaire, L. and P. Germain. 1863. Revision des Coléoptères du Chili, Suite (1). Fam. Scarabaeidae. Genres Maypa et Listronyx. Annales de la Société Entomologique de France, (4)2, 723-746. [Dated 1862].
Faldermann, F. 1835. Fauna entomologica trans-caucasia. Coleoptera: Bd. 1 Coleoptera persico-armeniaca. Nouveaux Mémoires de la Société Impériale des Naturalistes de Moscou 4: 1-310, 10 plates.
Fall, H. C. 1930. On Ataenius strigatus Say and allied species (Coleoptera). Journal of the New York Entomological Society 38: 93-108.
Fallén, C. F. 1807. Observationes Entomologicae, part 3. Litteris Berlingianis, Lund. Pp. 31-42.
Felsche, C. 1910. Über coprophage Scarabaeiden. Deutsche Entomologische Zeitschrift 1910: 339-352.
Frey, G. 1967. Die Gattung Plectris (Philochlaenia). (Coleoptera, Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey 18: 1-136.
Frey, G. 1969a. Betimmumgstabelle und Revision der Gattung Liogenys Guérrez. (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten aus dem Museum G. Frey 20: 36-64.
Frey, G. 1969b. Neue Macrodactylini (Col. Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey 20: 376-402.
Frey, G. 1970. Beitrag zur Kenntnis der südamerikanischen Melolonthiden (Col.). Entomologische Arbeiten aus dem Museum G. Frey 21: 281-287.
Frey, G. 1973. Synopsis der südamerikanischen Sericinen (Col., Scarab., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 24: 315-366.
Germain, P. 1897. Apuntes entomolójicos. Anales de la Universidad. Republica de Chile 98: 287-300.
Germain, P. 1901. Apuntes entomologicos. El jénero Phytoloema. Anales de la Universidad. Republica de Chile 108: 1-21.
Germain, P. 1903a. Apuntes entomológicos. Anales de la Universidad. Republica de Chile, 112-113, 391-445.
Germain, P. 1903b. Lista de los insectos traidos por los Señores Reiche i Machado de un viaje á Lebu y á La Mocha. Anales del Museo Nacional de Chile 16: 49-63.
Germain, P. 1905. Apuntes entomológicos. Anales de la Universidad. Republica de Chile 115: 449-506. [Dated 1904].
Germain, P. 1911. Catálogo de los coleópteros chilenos del Museo Nacional. Boletin del Museo Nacional de Chile 3(1): 47-73.
Germain, P. and R. A. Philippi. 1854. Description de 21 especies nuevas de Coleópteros de Chile. Preceden algunas observaciones jenerales sobre los Coleópteros de Chile. Anales de la Universidad de Chile 11: 325-336.
Germar, E. F. 1824. Insectorum Species Novae aut Minus Cognitae, Descriptionibus Illustratae. Coleoptera, volume 1. J. C. Hendelii et Filii, Halle. 624 pp.
Germar, E. F. 1843. Ueber die Gruppe der kugelförmigen Trogiden. Zeitschrift für die Entomologie 4: 109-148.
Germar, E. F. 1848. Beiträge zur Insektenfauna von Adelaide. Linnaea Entomologica 3: 153-247.
Gillet, J. J. E. 1911. Lamellicornes coprophages nouveaux ou peu connus d'Amérique du sud. Annales de la Société Entomologique de Belgique 55: 315-319.
Gmelin, J.F. 1790. Caroli a Linné Systema Naturae (13th edition), volume 1, part 4. G. E. Beer, Leipzig, pp. 1517-2224.
Gordon, R. D. and R. Pittino. 1992. Current status of the American genera and species of Psammodiini (Coleoptera: Scarabaeidae: Aphodiinae). Coleopterists Bulletin 46: 260-273.
Guérin-Méneville, F.E. 1830. Iconographie du Règne Animal de G. Cuvier, ou Représentation d'après Nature de L'une des Espèces les plus Remarquables et souvent non encore Figurees, de Chaque Genre d'Animaux, vol. 3. J. B. Baillière, Paris. Plates 21-24.
Guérin-Méneville, F. E. 1831. Insectes. In M. L. I. Duperrey (ed.), Voyage Autour du Monde, Exécuté par Ordre du Roi, sur la Corvette de la Majesté, la Coquille, Pendant les Années 1822, 1823, 1824 et 1825, sous le Ministère et Conformément aux Instructions de S. E. M. Le Marquis de Clermont-Tonnerre, Ministre de la Marine; et Publié sous les Auspices de son Excellence Mgr le Cte de Chabrol, Ministre de la Marine et des Colonies, Atlas (Histoire naturelle, Zoologie), Arthus Bertrand, Paris, 21 plates.
Guérin-Méneville, F. E. 1838a. Insectes du voyage de la Favorite. Magasin de Zoologie 8(9): 1-80.
Guérin-Méneville, F. E. 1838b. Insectes. Chapter 13, pp. 57-302. In M. L. I. Duperrey (ed.), Voyage Autour du Monde, Exécuté par Ordre du Roi, sur la Corvette de la Majesté, la Coquille, Pendant les Années 1822, 1823, 1824 et 1825, sous le Ministère et Conformément aux Instructions de S. E. M. Le Marquis de Clermont-Tonnerre, Ministre de la Marine; et Publié sous les Auspices de son Excellence Mgr le Cte de Chabrol, Ministre de la Marine et des Colonies, Volume 2, Part 2 (Zoologie, M. Lesson ed.), Division 1. Arthus Bertrand, Paris, 319 pp. ["1830"].
Guérin-Méneville, F. E. 1839. Description de quelques Coléoptères des côtes du détroit de Magellan. Revue Zoologique 1839:295-305.
Guérin-Méneville, F. E. 1844. Iconographie du Règne Animal de G. Cuvier, ou Représentation d'après Nature de L'une des Espèces les plus Remarquables et souvent non encore Figurees, de Chaque Genre d'Animaux, vol. 3. J. B. Baillière, Paris. 576 pp.
Guérin-Méneville, F. E. 1845. [Description of Ptilophyllum]. Revue Zoologique 1845: 439.
Gutiérrez, R. 1940a. Contribuciones al estudio de los Scarabaeidae Chilenos. Revista Chilena de Historia Natural 44: 93-99.
Gutiérrez, R. 1940b. Contribuciones al estudio de los Scarabaeidae Chilenos la subfamilia Scarabaeinae. Revista Chilena de Historia Natural 44: 275-280.
Gutiérrez, R. 1943. Contribuciones al conocimiento de los Scarabaeidae chilenos: la subfamilia Glaphyrinae. Revista Chilena de Historia Natural 45: 146-154. [Dated 1941].
Gutiérrez, R. 1944. Contribuciones al conocimiento de los Scarabaeidae chilenos: la tribu Liparetrini (Melolonthinae). Revista Chilena de Historia Natural 46: 117-131. [Dated 1942].
Gutiérrez, R. 1945. Contribuciones al conocimiento de los Scarabaeidae chilenos. Dynastinae Oryctini. Revista Chilena de Historia Natural 48: 112-118. [Dated 1944].
Gutiérrez, R. 1946. Notas sobre Scarabaeidae Chilenos. Livro homenagem R.F. d'Almeida 2: 11-16.
Gutiérrez, R. 1947. Escarabajos comunes a Chile y la Argentina (Col. Scarabaeidae). Revista de la Sociedad Entomológica Argentina 13:309-314.
Gutiérrez, R. 1949. Notas sobre Scarabaeidae neotrópicos (Coleoptera Lamellicornia). Anales de la Sociedad Científica Argentina, 148(6), 9-35.
Gutiérrez, R. 1950a. Notas sobre Scarabaeidae Chilenos (Coleoptera Lamellicornia). Arthropoda 1(2/4): 267-278.
Gutiérrez, R. 1950b. Scarabaeidae del Norte de Chile (Coleopt. Lamellic.). Anales de la Sociedad Científica Argentina 149(2): 52-75.
Gutiérrez, R. 1951. El genero Liogenys en Chile. Revista Chilena de Entomologia 1: 129-145.
Gutiérrez, R. 1952. Notas sobre Scarabaeidae neotrópicos (III). Revista Chilena de Entomologia 2: 207-227.
Haldeman, S. S. 1846. On several new genera and species of insects. Proceedings of the Academy of Natural Sciences of Philadelphia 3: 124-128.
Harold, E. 1859. Beiträge zur Kenntniss einiger coprophagen Lamellicornien (Erstes Strück). Berliner Entomologische Zeitschrift 3: 193-224.
Harold, E. 1862. Beitrage zur Kenntniss einiger coprophagen Lamellicornien (Viertes Stück). Berliner Entomologische Zeitschrift 6: 379-403.
Harold, E. 1867. Diagnosen neuer Coprophagen. Coleopterologische Hefte 1: 76-83.
Harold, E. 1868a. Die chilensischen Aphodiden. Berliner Entomologische Zeitschrift 11: 278-282. [1867].
Harold, E. 1868b. Diagnosen neuer Coprophagen. Coleopterologische Hefte 3: 80-86.
Harold, E. 1868c. Diagnosen neuer Coprophagen. Coleopterologische Hefte 4: 79-86.
Harold, E. 1869a. Scarabaeidae. In M. Gemminger and E. Harold (eds.), Catalogus Coleopterorum Hucusque Descriptorum Synonymicus et Systematicus, vol. 4. E. H. Gummi, Munich. Pp. 979-1346.
Harold, E. 1869b. Diagnosen neuer Coprophagen. Coleopterologische Hefte 5: 95-104.
Harold, E. 1869c. Abänderungen vergebener Namen. Coleopterologische Hefte 5: 122-125.
Harold, E. 1871. Diagnosen neuer Coprophagen. Coleopterologische Hefte 8: 114-116.
Harold, E. 1872. Monographie der Gattung Trox. Coleopterologische Hefte 9-10: 1-192.
Harold, E. 1874. Beiträge zur Kenntniss einiger coprophagen Lamellicornien. (Achtes Stück). Berliner Entomologische Zeitschrift 18: 177-208.
Harris, T. W. 1827. Minutes towards a history of some American species of Melolonthae particularly injurious to vegetation. Massachusetts Agricultural Journal 10(1): 1-12.
Hawkins, S. J. 2006. A revision of the Chilean tribe Lichniini Burmeister, 1844 (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 1266: 1-63.
Heer, O. Fauna Coleopterorum Helvetica, part 1, fascicle 3. Orelii, Fuesslini et Sociorum, Zurich. Pp. 361-652.
Heller, K. M. 1908. Ein neuer Aclopide aus Argentinien (Col.). Deutsche Entomologische Zeitschrift 1908: 55-58.
Hinton, H. E. 1937. Descriptions of new American Ataenius, with notes on others (Coleoptera, Scarabaeidae). Annals and Magazine of Natural History series 10, volume 20: 177-196.
Hoffmannsegg, J. C. 1817. Entomologische Bemerkungen bei Gelegenheit der Abhandlungen über amerikanische Insecten. Zoologisches Magazin 1: 8-56.
Höhne, W. 1922. Aspidolea (subg. Aspidolites) atricollis n. sp. (Col. Dyn.). Deutsche Entomologische Zeitschrift 1922: 374-376.
Hope, F. W. 1837. The Coleopterist's Manual containing the lamellicorn insects of Linnaeus and Fabricius. London. 121 pp.
Hope, F.W. 1841. Descriptions of some nondescript Lamellicorn beetles in his collection. Annals and Magazine of Natural History series 1, volume 8: 302-304.
Hope, F. W. 1847. Descriptions of various coleopterous insects from New Holland, collected chiefly by Mr. Fortnum, at Adelaide. Transactions of the Entomological Society of London 4: 280-285.
Hopping, R. 1931. New Coleoptera from western Canada. Canadian Entomologist 63: 233-238.
Horn, G. H. 1887. A monograph of the Aphodiini inhabiting the United States. Transactions of the American Entomological Society 14: 1-110.
Howden, H. F. and B. D. Gill. 2000. Tribes of New World Ceratocanthinae, with keys to genera and descriptions of new species (Coleoptera: Scarabaeidae). Sociobiology 35(2B): 281-329.
Jameson, M. L. and A. B. T. Smith. 2002. Revision of the South American genus Brachysternus Guérin-Méneville (Coleoptera: Scarabaeidae: Rutelinae: Anoplognathini: Brachysternina). Coleopterists Bulletin 56: 321-366.
Joly, L. J. and H. Escalona. 2002. Dos nuevas especies de Dyscinetus Harold de Argentina (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini). Entomotropica 17(3): 197-205.
Kirby, W. 1819. A description of several new species of insects collected in New Holland by Robert Brown. Transactions of the Linnean Society of London 12: 454-482. [Dated 1818.]
Kirby, W. 1837. The Insects. In J. Richardson, W. Swainson, and W. Kirby (eds.), Fauna Boreali-Americana, part 4. Josiah Fletcher, Norwich. 325 pp.
Klug, J. C. F. 1845. Coleoptera XV-XVI, 8 unnumbered pages, plate XLI-XLII. In C. G. Eherenberg (ed.), Symbolae Physicae seu Icones et Descriptiones Insectorum quae ex itinere Africam Borealem et Asiam Occidentalem, volume 5. G. Reimeri, Berlin.
Kolbe, H. J. 1893. Beiträge zur Kenntniss der Mistkäfer, Lamellicornia onthophila. Stettiner Entomologische Zeitung 54: 188-202.
Kolbe, H. 1907. Coleopteren. Ergebnisse der Hamburger Magalhaensische Sammelreise 8: 1-125.
Kolbe, H. J. 1908. Dynastidae, Cetoniinae und Scarabaeidae. In: L. Schultze, Zoologische und Arthropologische Ergebnisse einer Forschungsreise im Westlichen und Zentralen Südafrika. Bd. 1. Denkschriften der Medizinisch-Naturwissenschaftlichen Gesellschaft zu Jena 13: 121-132.
Kriesche, R. 1919. Neue Chiasognathinen (Coleopt. Lucanid.). Archiv für Naturgeschichte (A) 83: 93-95. [Dated 1917].
Kriesche, R. 1922. Zur Kenntnis der Lucaniden. Stettiner Entomologische Zeitung 83: 115-137.
Kugelann, J. G. and J. K. W. Illiger. 1798. Verzeichniss der Käfer Preussens. Johann Jacob Gebauer, Halle. 510 pp.
Lacordaire, J. T. 1856. Histoire Naturelle des Insectes. Genera des Coléoptères, vol. 3. Librairie Encyclopédique de Roret, Paris. 594 pp.
Lacroix, J. P. 1979. Contributions a l'etude des Coleopteres lucanides du globe: deux genres nouveaux et onze especes inedites (Chiasognathinae, Lucaninae, Chalcodinae, Cladognathinae, Dorcinae). Bulletin and Annales de la Société Royal Belge d'Entomologie 114: 249-294. [Dated 1978].
Landin, B.-O. 1955. Reports of the Lund University Chile Expeditions 1948-49. 22. Coleoptera; Lamellicornia. Lunds Universitets Arsskrift N. F. Avd 2. Bd 51. Nr 14: 1-14.
van Laicharting, J. N. 1781. Verzeichniss und Beschreibung der Tyrolen Insecten. 1. Theil Käferartige Insecten, volume 1. Johann Caspar Füessly, Zürich. 248 pp.
van Lansberge, J. W. 1874. Ennearabdus, gen. nov.; Synapsis Ritsemae Lansb. Annales de la Société Entomologique de Belgique 17: CXLIII-CXLIV.
van Lansberge, J. W. 1885. Descriptions d'espèces nouvelles de coléoptères appartenant au Musée Civique de Gênes. Annali del Museo Civico di Storia Naturale di Genova (2)2: 375-400.
Laporte, F. L. (Comte de Castelnau). 1840. Histoire Naturelle des Insectes Coléoptères (Volume 2 of Histoire Naturelle des Animaux Articulés). P. Duménil, Paris. 564 pp.
Latreille, P. A. 1797. Précis des Caractéres Génériques des Insectes, Disposés dans un Ordre Naturel. F. Bourdeaux, Paris, 201 pp.
Latreille, P. A. 1802. Histoire Naturelle, Genérale et Particuliere, des Crustacés et des Insectes, volume 3. F. Dufart, Paris, 467 pp.
Latreille, P. A. 1804. Histoire Naturelle, Genérale et Particuliere, des Crustacés et des Insectes, volume 10. F. Dufart, Paris, 445 pp.
Latreille, P. A. 1807. Genera Crustaceorum et insectorum Secundum Ordinem Naturalem in Familias Disposita, Iconibus Exemplisque Plurimis Explicata, volume 2. Amand Koenig, Paris, 280 pp.
Latreille, P. A. 1829. Le Régne Animal Distribué a'Après son Organisation, pour Servir de Base a l'Histoire Naturelle des Animaux et d'Introduction a l'Anatomie Comparée. Nouvelle Édition, Revue et Augmenté. Tome IV. Crustacés, Arachnides et Partie des Insectes. Déterville and Crochard, Paris. 584 pp.
Lea, A. M. 1923. On Australian aphodiides (Coleoptera). Proceedings of the Royal Society of Victoria 36(1): 1-20.
LeConte, J. L. 1857. Report upon insects collected on the survey. In, Reports of Explorations and Surveys for a Railroad Route from the Mississippi River to the Pacific Ocean, volume 12, part 3, pp. 1-72, 2 plates.
LePeletier, A. L. M. and J. G. A. Serville. 1828a. Scarabé, Scarabaeus (pp. 346-382). In, P. A. Latreille, A. L. M. LePeletier, J. G. A. Serville, and F. E. Guérin-Méneville (eds.) Encyclopédie Méthodique. Histoire Naturelle. Entomologie, ou Histoire Naturelle des Crustaces, des Arachnides et des Insectes, volume 10, part 2. Mme veuve Agasse, Paris, pp. 345-832.
Lewis, G. 1895. On the Lamellicorn Coleoptera of Japan, and notices of others. Annals and Magazine of Natural History (6) 16: 374-408.
Linell, M. L. 1896. New species of North American Coleoptera of the family Scarabaeidae. Proceedings of the United States National Museum 18: 721-731.
Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis, edition 10, volume 1. Laurentii Salvii, Stockholm. 823 pp.
Linnaeus, C. 1767. Systema Naturae, per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis, edition 12, volume 1, part 2. Laurentii Salvii, Stockholm. Pp. 533-1327
Luederwaldt, H. 1925. Novas especies do genero Pinotus (Col. Lamellic. – Coprinae). Boletim do Museu Nacional do Rio de Janeiro 2(1): 67-69.
MacLeay, W. S. 1819. Horæ Entomologicæ: or Essays on the Annulose Animals, volume 1, part 1. S. Bagster, London. 160 pp.
Maes, J.-M. 1992. Lista de los Lucanidae (Coleoptera) del mundo. Revista Nicaraguense de Entomología 22A: 1-60.
Mannerheim, C. G. 1829. Description de quarante nouvelles espèces de scarabéides du Brésil. Mémoires de la Société Impériale des Naturalistes de Moscou 7: 31-80, plate 1-2.
Marsham, T. 1802. Entomologia Britannica, Sistens Insecta Britanniae Indigena, Secundum Methodum Linnaeanam Disposita. Coleoptera, volume 1. Wilks and Taylor, London. 547 pp.
Martínez, A. 1944. Insectos nuevos o poco conocidos – II. Revista Argentina de Entomología 2: 34-40.
Martínez, A. 1945. Insectos nuevos o poco conocidos (IV) (Col. Scarabaeidae). Revista de la Sociedad Entomológica Argentina 12(5): 394-409
Martínez, A. 1950. Contribucion al conocimiento del genero "Megathopa" Eschsch., 1822, en la Argentina. Revista Española de Entomología 26: 197-269.
Martínez, A. 1952. Scarabaeidae nuevos o poco conocidos, III. Misión de Estudios de Patalogía Regional Argentina 23: 53-118.
Martínez, A. 1953a. Dos géneros nuevos de Melolonthinae (Coleopt. Scarabaeid.). Boletín de la Sociedad Entomológica Argentina 2: 3.
Martínez, A. 1953b. Un Nuevo género y especie de Canthonina (Coleopt. Scarabaeid.). Boletín de la Sociedad Entomológica Argentina 2: 3.
Martínez, A. 1954a. Nuevas especies de Oogenius Solier; Coleoptera, Scarabaeidae, Rutelinae. Revista Chilena de Entomología 3: 75-86. [Dated 1953].
Martínez, A. 1954b. Un nuevo género de Bolboceratini Neotropical (Coleoptera, Scarabaeidae). Revista Brasilera de Entomologia 2, 193-204.
Martínez, A. 1956. Scarabaeoidea neotropica. III. Una nueva especie de Sericoides (Col. Melolonth.). Neotropica 2(8): 45-48.
Martínez, A. 1957. Nuevos generos y especies de Macrodactylini (Col. Scarab. Melolonthinae). Neotropica 3(11): 47-52.
Martínez, A. 1959a. Scarabaeoidea neotropica. VI. Algunos Melolonthidae nuevos o poco conocidos (Col.). Neotropica 5(16): 23-29.
Martínez, A. 1959b. Catalogo de los Scarabaeidae Argentinos (Coleoptera). Revista del Museo Argentino de Ciencias Naturales "Bernardo Rivadavia" 5(1): 1-126, 4 plates.
Martínez, A. 1965. Insectos nuevo poco conocido. XII. Una nueva especies de Sericoides Guérin (Col. Scarab., Sericinae). Revista Sociedad Entomologica Argentina 27: 89-92. [Dated 1964].
Martínez, A. 1967. El género Athlia Erichson (Col. Scarabaeidae, Sericinae). Entomologische Arbeiten aus dem Museum G. Frey 18: 327-372.
Martínez, A. 1971. Un nueva especie de Sericoides Guérin de la Argentina (Col. Scarab. Sericinae, Liparetrini). Revista Sociedad Entomologica, Argentina 33: 115-120.
Martínez, A. 1972a. Un nuevo genero y especie de Pachydemini Chileno (Col. Scarabaeidae, Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey 23: 86-92.
Martínez, A. 1972b. Algunas consideraciones sobre los generos Apterodema Fairmaire y Sericoides Guerin, con descripcion de dos nuevas especies (Scarab. Sericinae, Liparetrini). Comunicaciones del Museo Argentino de Ciencias Naturales Bernardino Rivadavia 1(3): 37-62.
Martínez, A. 1973. El genero Neogutierrezia Martinez, 1953 (Col. Scarab. Melolonth. Pachydemini). Anales de la Sociedad Científica Argentina 195 (1-2): 25-41.
Martínez, A. 1974. Nuevas consideraciones sobre el Genero Athlia Erichson, 1835 (Col., Scarabaeidae, Sericinae). Entomologische Arbeiten aus dem Museum G. Frey 25: 343-353.
Martínez, A. 1975. Contribución al conocimiento de los Pachydemini neotropicales (Col. Scarabaeidae, Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey 26: 227-251.
Martínez, A. 1976. Contribucion al conocimiento de los Bolboceratini sudamericanos (Coleoptera, Scarabaeidae, Geotrupinae, Bolboceratini). Studia Entomologica 19: 531-551.
Martínez, A. 1981. Un Nuevo Ceratognathus neotropical (Coleoptera: Lucanidae). Boletín de Entomología Venezolana 2(2): 13-19.
Martínez, A. 1982. Un nuevo genero de Pachydemini Argentino (Col. Scarab. Melolonthinae). Revista Sociedad Entomologica Argentina 41(1-4): 89-91.
Martínez, A., F. S. Pereira, and M. A. Vulcano. 1970. Aegialiini, nueva tribu de Aphodiinae (Col. Scarabaeidae) para la region Neotropical. Acta Zoologica Lilloana 25(26): 335-341.
Melsheimer, F. E. 1846. Descriptions of new species of Coleoptera of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia 2: 134-160.
Ménétries, E. 1832. Catalogue Raisonné des Objets de Zoologie Recueillis dans un Voyage au Caucase et Jusqu'aux Frontières Actuelles de la Perse Entrepris par Ordre de S. M. l'Empereur. L'Académie Impériale des Sciences, St. Petersburg. 271 pp.
Molino-Olmedo, F. 2001. Cladistica de genero Chiasognathus Stephens, 1831 y Sphaenognathus Buquet, 1838 con proposicion de subgeneros nuevos (Coleoptera: Lucanidae). Revista Chilena de Entomología 28: 79-85.
Molino-Olmedo, F. 2002. Cladística del género Chiasognathus Stephens, 1831 con proposición de subgéneros nuevos (Coleoptera: Lucanidae). Elytron 15: 119-128. [Dated 2001].
Molino-Olmedo, F. 2003. Nombres de reemplazo para los subgéneros Ramirezia Molino-Olmedo, 2001 y Carmenia Molino-Olmedo, 2001 del género Chiasognathus Stephens, 1831 (Coleoptera: Lucanidae). Elytron 16: 29-30. [Dated 2002].
Mondaca, J. 2005. Utilización de Oogenius Solier, 1851 nuevo nombre para Oogenius gutierrezi Martínez y Peña, 1994 (Coleoptera: Scarabaeidae) de Chile, incluido listado de especies. Revista Chilena de Entomología 30(2): 17-20. [Dated 2004].
Mondaca, J. and M. J. Paulsen. 2005. Ceratognathus argentinensis Martínez, nuevo registro de Aesalinae (Coleoptera: Lucanidae) para Chile. Revista Chilena de Entomología 30(2): 21-26. [Dated 2004].
Montrouzier, X. Essai sur la faune entomologique de la Nouvelle-Calédonie (Balade) et de Iles des Pins, Art, Lifu, etc. Annales de la Société Entomologique de France 3(8): 229-308, plate 7.
Moser, J. 1918b. Neue Amerikanische Melolonthiden (Col.). Stettiner Entomologische Zeitung 79: 95-167.
Moser, J. 1918d. Beitrag zur Kenntnis der Melolonthiden (Col.). (VIII). Stettiner Entomologische Zeitung 79: 297-349.
Moser, J. 1919a. Beitrag zur Kenntnis der Melolonthiden (Col.). (VII. Fortsetzung.). Deutsche Entomologische Zeitschrift, 1919, pp. 1-34.
Moser, J. 1919c. Beitrag zur Kenntnis der Melolonthiden. (Col.) (X.). Stettiner Entomologische Zeitung, 80, 330-364.
Moser, J. 1921b. Neue Melolonthiden Mittel- und Süd-Amerika. Stettiner Entomologische Zeitung, 82, 133-182.
Moser, J. 1924a. Beitrag zur Kenntnis der Melolonthiden (Col.). (XIII). Stettiner Entomologische Zeitung, 84, 114-136.
Moser, J. 1924b. Beitrag zur Kenntnis der Melolonthiden (Col.). (XIV). Stettiner Entomologische Zeitung, 84, 137-164.
Moser, J. 1926. Beitrag zur Kenntnis der Melolonthiden (Col.). (XIV). Stettiner Entomologische Zeitung, 87, 198-208.
Motschulsky, V. 1859. Sur les collections coléoptèrologiques de Linné et de Fabricius (continuation). XXII. Lamellicornes. Etudes Entomologiques 8: 147-162.
Mulsant, E. 1842. Histoire Naturelle des Coléoptères de France. Lamellicornes. Maison, Libraire, Paris, 623 pp.
Mulsant, E. and C. Rey. 1870a. Description de quelques nouvelles espèces d'aphodiens (Colèoptères Lamellicornes). Opuscules Entomologiques 14: 203-221.
Mulsant, E. and C. Rey. 1870b. Tribu des Lamellicornes. Annales de la Société d'Agriculture, Sciences et Industries de Lyon Série 4, Tome 2: 241-650.
Nagel, P. 1928. Neues über Hirschkäfer (Col. Lucan.). Deutsche Entomologische Zeitschrift 1928: 273-298.
Nagel, P. 1932. Neues über Hirschkäfer. Entomologische Blätter 28: 113-121.
Nonfried, A. F. 1894. Beschreibungen neuer Lamellicornier, Buprestiden und Cerambyciden aus Central- und Süd-Amerika. Entomologische Nachrichten 20:113-128.
Numhauser, J. 1985. Revision del genero Pycnosiphorus grupo I lessoni Weinreich (Coleopteros. Lucanidae). Revista Chilena de Entomologia 12: 53-75.
Ocampo, F. C. 2004. Food relocation behavior and synopsis of the southern South American genus Glyphoderus Westwood (Scarabaeidae: Scarabaeinae: Eucraniini). Coleopterists Bulletin 58: 295-305.
Ohaus, F. 1904a. Revision der Anoplognathiden (Coleoptera lamellicornia). Stettiner Entomologische Zeitung 65:57-175.
Ohaus, F. 1904b. Revision der amerikanischen Anoplognathiden (Coleoptera lamellicornia). Stettiner Entomologische Zeitung 65:254-341.
Ohaus, F. 1905a. Revision der amerikanischen Anoplognathiden (Coleoptera lamellicornia). Stettiner Entomologische Zeitung 66:120-167.
Ohaus, F. 1905b. Beiträge zur Kenntniss der amerikanischen Ruteliden. Stettiner Entomologische Zeitung 66: 283-329.
Ohaus, F. 1909a. Neue Coleoptera lamellicornia aus Argentinien. Deutsch Entomologische Zeitschrift 1909: 425-447.
Ohaus, F. 1909b. Nachträge und Berichtigungen zu meiner Revision der Brachysterniden (Coleopt. lamellicornia). Stettiner Entomologische Zeitung 71:3-26. ["1910"].
Ohaus, F. 1910. Neue Coleoptera lamellicornia aus Argentinien. Deutsch Entomologische Zeitschrift 1910: 173-186.
Ohaus, F. 1911a. Neue Coleoptera lamellicornia aus Argentinien (Col.). Deutsch Entomologische Zeitschrift 1911: 553-565.
Ohaus, F. 1911b. Isoceraspis, subgen nov. melolonthinarum (Col.). Deutsche Entomologische Zeitschrift, 1911, pp. 669-670.
Ohaus, F. 1918. Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae. Coleopterorum Catalogus 20:1-241. ["1915"].
Olivier, A. G. 1789a. Coléoptères, part 1. Number 3, Scarabé, Scarabaeus, 190 pp. In Entomologie, ou Histoire Naturelle des Insectes, avec leurs Caractères Génériques et Spécifiques, leur Description, leur Synonymie, et leur Figure Enluminée. Baudouin, Paris.
Olivier, A. G. 1789b. Coléoptères, part 1. Number 4, Trox, Trox, 14 pp. In Entomologie, ou Histoire Naturelle des Insectes, avec leurs Caractères Génériques et Spécifiques, leur Description, leur Synonymie, et leur Figure Enluminée. Baudouin, Paris.
d'Orbigny, H. 1896. Synopsis des aphodiens d'Europe et du basin de la Méditerranèe. L'Abeille Journal d'Entomologie 18: 197-271.
Özdikmen, H. and S. Turgut. 2006. Nomenclatural changes in Lucaninae (Coleoptera: Lucanidae). Munis Entomology and Zoology 1(1): 145-148.
Panzer, G. W. F. 1796. Scarabaeus caesus. Der Dungkäfer mit der Brustschramme. Faunae Insectorum Germanicae initia oder Deutschlands Insecten 35: 2.
Parry, F. J. S. 1864. A catalogue of lucanoid Coleoptera; with illustrations and descriptions of various new and interesting species. Transactions of the Entomological Society of London, series 3, volume 2:1-113, plates 1-12.
Parry, F. L. S. 1870. A revised catalogue of the lucanoid Coleoptera; with remarks on the nomenclature, and descriptions of new species. Transactions of the Entomological Society of London 1870: 53-118, plates 1-3.
Paulian, R. 1934. Deux nouveaux coprophages des Nouvelles-Hébrides. Revue Franaise d'Entomologie 1: 219-221.
Paulian, R. 1937. Trois nouveaux Ataenius de la région australienne. Miscellanea Entomologica 38: 41-43.
Paulian, R. 1979. Un nouvel Aphodiidae du Portugal (Col.). Bulletin de la Société Entomologique de France 84: 66-67.
Paulian, R. 1982. Révision des Cératocanthides (Coleoptera Scarabaeoidea) d'Amérique du Sud. Mémoires du Muséum National d'Histoire Naturelle, Série A, Zoologie 124: 1-110 + XVIII plates.
Paulian, R. 1984. Les Orphnidae Américains (Coléoptères, Scarabaeoidea). Annales de la Société Entomologique de France 20: 65-92.
Paulsen, M. J. 2005. A revision of the southern South American stag beetles of the genus Sclerostomus Burmeister (Coleoptera: Scarabaeoidea: Lucanidae). Zootaxa 1060:1-26.
Paulsen, M. J. and J. Mondaca. 2006. Revision of the South American Ceratognathini (Coleoptera: Lucanidae: Aesalinae) with the description of a new genus and a new species. Zootaxa 1191: 1-19.
von Paykull, G. 1798. Fauna Suecica. Insecta, volume 1. Joh. F. Edman, Uppsala. 358 pp.
Pereira, F. S. and A. and Martínez. 1960. Tres nuevos generos de Canthonini Americanos (Coleoptera, Scarabaeidae). Acta Zoologica Lilloana 17: 165-184.
Péringuey, L. 1901. Descriptive catalogue of the Coleoptera of South Africa. Fam. Lucanidae. Fam. Scarabaeidae. Sub-families: Coprinae, Aphodiinae, Troginae, Chironinae, Geotrupinae, Hybosorinae, Orphninae, Dynastinae. Transactions of the South African Philosophical Society 12: 1-560.
Petrovitz, R. 1963. Neue und verkannte Aphodiinae aus allen Erdteilen (Col. Scarab.). Entomologische Arbeiten aus dem Museum G. Frey 14: 630-647.
Petrovitz, R. 1971. Neue Aphodiidae der äthiopischen und australischen Region (Col.). Entomologische Arbeiten aus dem Museum G. Frey 22: 99-104.
Petrovitz, R. 1973. Neue Aphodiinae, Hybosoridae und Acanthocerinae aus der Neotropischen Region (Coleoptera: Scarabaeidae). Studia Entomologica 16: 141-202.
Philippi, F. 1859. Algunas especies nuevas de Coleópteros de la provincia de Valdivia. Anales de la Universidad de Chile 16: 656-678.
Philippi, F. 1861. Observaciones sobre los lamelicornios de Chile, descritos en la obra del Señor Gay, con descripcion de algunas especies nuevas. Anales de la Universidad, Republica de Chile 18:735-742.
Philippi, F. (1887) Catálogo de los Coléopteros de Chile. Anales de la Universidad Republica de Chile, 71, 619-806.
Philippi, R.A. (1864) Sobre algunos Coléopteros nuevos de Chile de la familia de las melolontideas. Anales de la Universidad Republica de Chile, 24, 434-462.
Philippi, R. A. 1866. Einige Insekten von Chile. Stettiner Entomologische Zeitung 27: 109-117.
Philippi, R. A. 1873. Chilenische Insekten. Entomologische Zeitung 34: 296-316, plate 1-2.
Philippi, R. A. and F. Philippi. 1864. Beschreibung einiger neuen Chilenischen Käfer. Stettiner Entomologische Zeitung 25:313-406.
Pittino, R. 1987. New Coleoptera Trogidae from South America (XXXII contribution to the knowledge of Coleoptera Scarabaeoidea). Giornale Italiano di Entomologia 3(17): 377-397.
Prell, H. 1936. Beiträge zur Kenntnis der Dynastinen. Über die Homonymieverhältnisse der Namen von Gattungen und Untergattungen. Entomologische Blätter 32: 145-152
Rakovic, M. 1981a. Revision of species of the tribe Psammodiini from the Australian region (Coleoptera, Scarabaeidae, Aphodiinae) I. key to genera and revision of the genera Aphodopsammobius Endrödi, Psammodius Fallén, Diastictus Mulsant, Phycochus Broun and Tesarius g. n. Annotationes Zoologicae et Botanicae 139: 1-37.
Rakovic, M. 1981b. A revision of the Psammodius Fallen species from Europe, Asia and Africa. Rozpravy Ceskoslovenske Akademie Ved 91: 1-82.
Rakovic, M. 1990. Review of the genus Leiopsammodius Rakovic on the world basis with a key to species from the western hemisphere and description of a new species (Coleoptera, Scarabaeidae, Aphodiinae). Annotationes Zoologicae et Botanicae 197: 1-18.
Redtenbacher, L. 1868. Reise der österreichen Fregatte Novara um die Erde in der Jahren 1857, 1858, 1859 unter der befehlen des Commodore B. Wüllerstorf-Urbair. Zoologische Theil. Zweiter Band: Coelopteren. Wien. 249 pp.
Reiche, L. 1841. Tableau d'une division systématique de la tribu des coprophages, dans la famille des Lamellicornes. Revue Zoologique 1841: 211-213.
Reiche, L. 1850a. La description suivante d'une nouvelle espèce de Chiasognathus. Revue et Magasin de Zoologie 2(2): 249-250.
Reiche, L. 1850b. Mémoire sur les genres Psalidognathus et Chiasognathus. Annales de la Société Entomologique de France 2(8): 263-269.
Reitter, E. 1892. Bestimmungs-Tabelle der Lucaniden und coprophagen Lamellicornen des palaearctischen Faunengebietes. Verhandlungen des Naturforschenden Vereins in Brünn 24: 1-230.
Richards, A. M. 1959. Revision of the genus Saprosites Redtenbacher, 1858 (Coleoptera: Aphodiinae) in New Zealand. Transactions of the Royal Society of New Zealand 87: 35-50.
Ripa, S. R., P. S. Rojas, and G. Velasco. 1995. Releases of biological control agents of insect pests on Easter Island (Pacific Ocean). Entomophaga 40 (3/4): 427-440.
Robinson, M. 1948. A new genus of Scarabaeidae (Coleoptera). Notulae naturae of the Academy of Natural Sciences of Philadelphia 211: 1-2.
Roig-Juñent, S., G. E. Flores, F. C. Ocampo, and A. B. T. Smith. 2005. Nuevas citas de Coleoptera para la Argentina (Carabidae, Lucanidae, Scarabaeidae y Tenebrionidae). Revista de la Sociedad Entomológica Argentina 63(3-4): 45-48.
Roth, J. R. 1851. Diagnosen neuer Coleoptera aus Abyssinien. Archiv für Naturgeschichte 17(1): 115-133.
Ruiz, F. 1924. El género Idiostoma en Chile. Revista Chilena de Historia Natural 28: 99-101.
Ruiz-Manzanos, E. 2006. Redescription and first record of Diaphylla granulata for Argentina (Coleoptera: Scarabaeidae: Melolonthinae). Revista de la Sociedad Entomológica Argentina 65(3-4): 79-86.
Say, T. 1835. Descriptions of new North American Coleopterous insects, and observations on some already described. Boston Journal of Natural History 1: 151-203.
Saylor, L.W. 1937a. Synopsis of the beetles of the Chilean genus Phytholaema (Scarabaeidae: Melolonthinae). Proceedings of the United States National Museum, 85: 5-11.
Saylor, L.W. 1945. Studies in the melolonthine scarab beetle genera of the American continents. No. IV. A new genus from Argentina. Revista de Entomologia 16(3): 441-444.
Saylor, L.W. (1946a) A new West Indian scarab beetle. Bulletin of the Brooklyn Entomological Society, 41(1), 1-3.
Saylor, L.W. (1946b) Studies in the melolonthine scarab beetle genera of the American continents. I. Revision of the genus Athlia. Proceedings of the Entomological Society of Washington, 48(1), 18-25.
Schmidt, A. 1907. Namensanderungen in der Gattung Aphodius und eine Neubescrheibung. Deutsche Entomologische Zeitschrift 1907: 201-202.
Schmidt, A. 1909a. Eine Serie neuer Aphodiinen und eine neue Gattung. Societas Entomologica 24(2): 10-12.
Schmidt, A. 1909b. Eine Serie neuer Aphodiinen und eine neue Gattung. Societas Entomologica 24(8): 60-62.
Schmidt, A. 1909c. Eine Serie neuer Aphodiinen und eine neue Gattung. Societas Entomologica 24(9): 66-67.
Schmidt, A. 1910. Neue Arten aus den Gattungen Aphodius Illig., Ataenius Har., Saprosites Redtenb. (Col.). Deutsche Entomologische Zeitschrift 1910: 353-361.
Schmidt, A. 1911. Eine Serie neuer Aphodiinen. Entomologische Zeitung 72: 3-51.
Schmidt, A. 1912. Neue Scarabaeiden aus den Subfamilien Aphodiinae, Orphinae und Hybosorinae. Archiv für Naturgeschichte 78(A): 196-202.
Schmidt, A. 1913. Erster Versuch einer Einteilung der exotischen Aphodien in Subgenera und als Anhang einige Neubeschreibungen. Archiv für Naturgeschichte 79(A): 117-178.
Schmidt, A. 1931. Coleoptera-Scarabaeidae-Aphodiinae von Juan Fernandez, p. 638. In Skottsberg, C. (ed.) The Natural History of Juan Fernandez and Easter Island, volume 3, Zoology. Almquist and Wiksells, Upsalla. 688 pp.
Scholtz, C. H. 1982. Catalogue of world Trogidae (Coleoptera: Scarabaeoidea). Entomology Memoir, Department of Agriculture and Fisheries, Republic of South Africa 54: 1-27.
Scholtz, C. H. 1990. Revision of the Trogidae of South America (Coleoptera: Scarabaeoidea). Journal of Natural History 24: 1391-1456.
Schreibers, C. F. A. 1802. Descriptions of some singular coleopterous insects. Transactions of the Linnean Society of London 6: 185-206, plate 19-21.
Schultze, W. 1916. Beitrage zur coleopteren Fauna der Philippinen. Philippine Journal of Science D11: 291-299, 2 plates.
Schweiger, H. 1959. Über einige von der Skottsbergexpedition im Antarkto-Archiplata-Gebiet aufgesammelte Koleopteren. Arkiv för Zoologi 12(1): 1-43.
Scopoli, J. A. 1763. Entomologia Carniolica Exhibens Insecta Carnioliae Indigena et Distributa in Ordines, Genera, Species, Varietates. Methodo Linnaeana. Trattner, Vienna. 420 pp.
Shipp, J. W. 1895. Notes on Onthophagus, Latr.; with corrections of nomenclature, and a description of a new genus. Entomologist 28: 178-179.
Skelley. P. E. 2001. New record for Tesarius Rakovic in South America (Coleoptera: Scarabaeidae: Aphodiinae: Psammodiini). Insecta Mundi 14: 240.
Smith, A. B. T. 2002. Revision of the southern South American endemic genus Aulacopalpus Guérin-Méneville with phylogenetic and biogeographic analyses of the subtribe Brachysternina (Coleoptera: Scarabaeidae: Rutelinae: Anoplognathini). Coleopterists Bulletin 56(3): 379-437.
Solier, A. J. J. 1851. Orden III. Coleopteros, pp. 5-285. In C. Gay (ed.), Historia Fisica y Politica de Chile. Zoología, vol. 5. C. Gay, Paris. 564 pp.
Stebnicka, Z. 2003. Revision and hypothetical phylogenetic analysis of the species of the New World genus Ataeniopsis (Coleoptera: Aphodiinae: Eupariini). European Journal of Entomology 100(1): 101-113.
Stebnicka, Z. 2004. The New World species of Ataenius Harold, 1867. IV. Revision of the A. strigicauda-group (Coleoptera: Scarabaeidae: Aphodiinae: Eupariini). Acta Zoologica Cracoviensia 47(3-4): 211-228.
Stebnicka, Z. T., M. Dellacasa, and P. E. Skelley. 2004. Review of the New World Aegialiini (Coleoptera: Scarabaeidae: Aphodiinae), with descriptions of two new genera from South America. Insecta Mundi 17:73-83. [Dated 2003].
Steinheil, E. 1874. Symbolae ad historian coleopterorum Argentiniae meridionalis, ossia enumeriazione dei coleotteri raccolti dal prof. P. Strobel nell'Argentinia meridionale, e descrizione dell species nuove, di Edoardo Stenheil. (Traduzione dal manoscritto tedesco inedito per P. Strobel, coll'aggiunta delle notizie sulla dimora delle specie.). Atti della Società Italiana di Scienze Naturali 15: 554-578. [1872].
Stephens, J. F. 1830. Illustrations of British Entomology; or, a Synopsis of Indigenous Insects: Containing their Generic and Specific Distinctions; with an Account of their Metamorphoses, Times of Appearance, Localities, Food, and Economy, as Far as Practicable. Mandibulata, volume 3. Baldwin and Cradock, London. 379 pp.
Stephens, J. F. 1831. Description of Chiasognathus grantii, a new lucanideous insect forming the type of an undescribed genus, together with some brief remarks upon its structure and affinities. Transactions of the Cambridge Philosophical Society 4(2): 209-217, plate IX-X.
Thomson, J. 1862. Catalogue des lucanides de la collection de M. James Thomson suivi d'un appendix renfermant la description des coupes génériques et spécifiques nouvelles. Annales de la Société Entomologique de France 4(2): 389-436.
Vaurie, P. 1962. A revision of the genus Trox in South America (Coleoptera, Scarabaeidae). Bulletin of the American Museum of Natural History 124: 101-167.
Vaz-de-Mello, F. Z. and G. Halffter. 2006. A new dung beetle genus with two new species from Chile (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 1193: 59-68.
Walker, F. 1858. Characters of some apparently undescribed Ceylon insects. Annals and Magazine of Natural History 2(3): 202-209.
Waterhouse, G. R. 1842. Description of a new species of lamellicorn beetle, brought from Valdivia by C. Darwin, Esq. Entomologist 1(18): 281-283.
Weinreich, E. 1958. Neue Arten der Gattung Pycnosiphorus (Col., Lucanidae) aus dem Senckenberg-Museum. Senckenbergiana Biologica 39(3/4): 169-176.
Westwood, J. O. 1837. Describing several new species of insects belonging to the family of the sacred beetles. Proceedings of the Zoological Society of London 5: 12-13.
Westwood, J. O. 1838a. Descriptions of several new species of insects belonging to the family of the sacred beetles. Transactions of the Zoological Society of London 2(2): 155-163.
Westwood, J. O. 1838b. Lucanidarum novarum exoticarum descriptions, cum monographia generum nigidii et figuli. Entomological Magazine 5(3): 259-268.
Westwood, J. O. 1845. A Catalogue of the Lucanoid Coleoptera, in the Collection of the Rev. F. W. Hope, M. A., F. R. S., &c. President of the Entomological Society of London. Together with Descriptions of the New Species Therein Contained. J. C. Bridgewater, London, 31 pp.
White, A. 1842. Descriptions of some hemipterous insects of the section Heteroptera. Transactions of the Entomological Society of London 3: 84-94.
Wiedermann, C. R. W. 1823. Zweihundert neue Käfer von Java, Bengalen und dem Vorgebirge der guten Hoffnung. Zoologisches Magazin 2(1): 3-133.
Wollaston, T. V. 1867. Coleoptera Hesperidium, being an enumeration of the coleopterous insects of the Cape Verde Archipelago. John Van Voorst, London, 285 pp.
Yawata, H. 1943. Eine neue Aphodius-Art aus Japan. Transactions of the Kansai Entomological Society 13: 7.
Zunino, M. 1984. Analisi sistematica e zoogeografica della sottofamiglia Taurocerastinae Germain (Coleoptera, Scarabaeidae: Geotrupidae). Bullettino del Museo Regionale di Scienze Naturali Torino 2(2): 445-464.
Zunino, M. and E. Barbero. 1991. Note su alcuni Scarabaeoidea (Coleoptera) della Terra del Fuoco, pp. 142-147. In B. Battaglia, P. M. Bisol, and V. Varotto (eds.), Atti del Primo Convegno di Biologia Antartica. Edizioni Universitarie Patavine, Padova. 408 pp.