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Classification
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........Heterogomphus Burmeister,
1847
........Heterogomphus Burmeister
1847: 224.
........Stypotrupes Burmeister 1847: 210 (synonym).
........Daemonoplus Thomson 1859a: 68 (subgenus).
........Baryxenus Bates 1891: 33 (synonym).
........Syneterogomphus Prell 1912a: 53 (synonym).
........Ortheterogomphus Prell 1912b: 103 (synonym).
........Heterogomphidium Prell 1912b: 105 (synonym).
........Hoplitogomphus Prell 1912b: 123 (synonym).
........Dineterogomphus Prell 1912b: 129 (synonym).
........Psileterogomphus Prell 1912b: 141 (synonym).
........Neobaryxenus Prell 1912b: 148 (synonym).
........Anoplogomphus Prell 1912b: 149 (synonym).
........Notheterogomphus Prell
1912b: 156 (synonym).
........Tracheterogomphus Prell
1912b: 161 (synonym). |
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Heterogomphus chevrolati.
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Distribution: Heterogomphus occurs
in Central and South America.
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Composition: The
genus Heterogomphus contains 47 species (Endrödi 1976b,
1985a; Dechambre 1986a).
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Diagnosis: Species in the genus Heterogomphus may
be recognized by the combination of quadridentate protibia, teeth of
the protibia directed obliquely forward, males with a large head horn,
females with one or two tubercles on the head (only one tubercle in the
species considered here; never with a horn), and prosternal process long
and stout.
Heterogomphus females,
especially H. mniszechi, are easily confused with the females
of some Golofa species (Dynastini). They are all about the same size,
black, and with densely punctate elytra. In H. mniszechi females,
the basal segment of the protarsus is distinctly shorter than the apical spur
of the protibia. In Golofa females, the basal segment of the protarsus
is subequal to or longer than the apical spur of the protibia (except for G. tersander Burmeister,
where it may occasionally be shorter). In addition, while the apex of the prosternal
process may have long, dense setae in both genera, the shaft of the process
is normally glabrous or sparsely setose in Heterogomphus and densely
setose in Golofa. Prell (1912b) was the first to provide a
comprehensive treatment of the genus, and Endrödi (1976b, 1985a)
gave the most recent synopsis.
The subgenus Daemonoplus contains those species with the apex
of the metatibia with a central tooth and the pronotum of the males with
three horns (H. mniszechi [Thomson] and H. flohri [Kolbe]). |
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Biological notes: In spite of the occasional abundance of some species
of Heterogomphus,
very little is known of their biology or immature stages. Adults are
nocturnal, and many are attracted to lights at night. Where the larvae
are known, they seem to feed on organic debris in the soil.
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Author: Brett
Ratcliffe
Generated on:26/MAR/2007.....Last
modified: 26/MAR/2007
University of Nebraska State Museum - Division of Entomology
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