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1.
.
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Mentum strongly
expanded, covering bases of labial palpi. Body
usually flattened |
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Phileurini
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1'.
.
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Mentum
narrow, not covering bases of labial palpi (Fig. 40, lower
figure). Body not flattened |
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2(1).
.
. |
Head and pronotum
lacking tubercles, horns, carinae, or fovea. Claw with onychium bisetose.
Male front claws in many species enlarged (larger than those in female).
Tarsi cylindrical |
.
.
Cyclocephalini |
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2'.
.
.
.
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Head
and/or pronotum (whether males or females) with
distinct tubercles, horns, carinae, or fovea. Claw with
onychium bisetose to multisetose. Male front claws enlarged or not.
Tarsi cylindrical to triangular |
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3(2).
.
. |
Posterior
tarsal segments (especially first or second) triangularly
expanded. Combined length of foretibia and foretarsus similar in both
sexes. Propygidium with or without stridulatory area |
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3'.
.
.
.
. |
Posterior tarsal
segments usually cylindrical, not triangularly
expanded (although basal segment with strong apical spine). Combined
length of foretibia and foretarsus longer in males than in females
(slightly less noticeable in Nearctic species). Propygidium usually
without distinct stridulatory area |
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4(3).
.
. |
Elytra covered
with dense tomentum (Spodistes, Lycomedes) or always
irregularly punctate, not with distinct rows of punctures. Body slightly
depressed |
.
. Agaocephalini |
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4'.
.
. |
Elytra
never with dense tomentum, punctures usually in distinct rows (except
for some densely punctate Heterogomphus species)
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5
(4).
. |
Apex of posterior
tibia usually truncate or smooth (Figs. 41-45). Sexual dimorphism
usually not well-pronounced |
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5'.
. |
Apex
of posterior tibia usually strongly crenulate or with
distinct teeth (Figs. 46-50). Sexual dimorphism usually distinct |
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Oryctini
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