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(Coleoptera: Scarabaeoidea)

M.J. Paulsen
(After Paulsen 2007)

Cucochodaeus sparsus (LeConte)
Photo by M.J. Paulsen.


Nearctic Ochodaeidae

CHAETOCANTHINAE, Scholtz in Scholtz et al.1988: 231
  Pseudochodaeus Carlson & Ritcher, 1801: 254 info  
    Pseudochodaeus estriatus Schaeffer, 1906: 271 (Ochodaeus) USA: CA, OR.
OCHODAEINAE Mulsant & Rey, 1871: 236
  Codocera Eschscholtz, 1821: 398 info
    syn. Stomphax Fischer von Waldheim, 1823: 158  
    Codocera gnatho (Fall in Fall & Cockerell, 1907: 247) (Ochodaeus) USA: CA, AZ, NM; northern Mexico.
      syn. Ochodaeus nimius Fall in Fall & Cockerell, 1907: 248
  Cucochodaeus Paulsen, 2007: 9 info  
    Cucochodaeus sparsus (LeConte, 1868: 51) (Ochodaeus) Western USA, northern Mexico.
       syn. Ochodaeus mandibularis Linell, 1896: 723
  Neochodaeus Nikolajev, 1995: 77
    Neochodaeus frontalis (LeConte, 1863: 76) Southeastern USA; northeastern Mexico.
       syn. Ochodaeus complex LeConte, 1868: 51
    Neochodaeus praesidii (Bates, 1887: 106) Southwestern USA; Mexico.
    Neochodaeus repandus (Fall, 1909: 36) Southwestern USA; northern Mexico.
    Neochodaeus striatus (LeConte, 1854: 222) Southwestern USA; northern Mexico.
  Parochodaeus Nikolajev, 1995: 77
    Parochodaeus biarmatus (LeConte, 1868: 51) Southwestern USA; northern Mexico.
    Parochodaeus californicus (Horn, 1895: 224) USA: CA.
    Parochodaeus howdeni (Carlson, 1975: 60) USA: TX; northern Mexico.
    Parochodaeus inarmatus (Schaeffer, 1906: 270) Southwestern USA; northern Mexico.
    Parochodaeus duplex (LeConte, 1868: 51) Southwestern and central USA; northern Mexico.
      syn. Ochodaeus kansanus Fall, 1909: 34  
    Parochodaeus pectoralis (LeConte, 1868: 51) Southwestern USA; northern Mexico.
    Parochodaeus peninsularis (Horn, 1895: 224) USA: southern CA; Mexico: Baja.
    Parochodaeus pixius Paulsen 2011: 2 USA: TX.
    Parochodaeus ritcheri (Carlson, 1975: 62) USA: TX; northern Mexico.
  Xenochodaeus Paulsen, 2007: 3
    Xenochodaeus americanus (Westwood, 1852: 66) (Ochodaeus) Central USA; south-central Canada.
      syn. Ochodaeus opacus LeConte, 1868: 51  
Xenochodaeus luscinus (Howden, 1968: 51) (Ochodaeus)
Canada; USA: Rockies.
    Xenochodaeus musculus (Say, 1835: 178) (Ochodaeus) Eastern US and southeastern Canada.
    Xenochodaeus planifrons (Schaeffer, 1906: 269) (Ochodaeus) Southwestern USA; Mexico.
    Xenochodaeus simplex (LeConte, 1854: 222) (Ochodaeus) Canada. USA: High Plains to southwest.
    Xenochodaeus ulkei (Horn, 1876: 182) (Ochodaeus) Northwest USA; Canada: BC.



Bates, H. W. 1887. Insecta. Coleoptera. Pectinicornia and Lamellicornia. Biologia Centrali-Americana 2(2): 1-432.

Carlson, D. C. 1975. Taxonomic characters of the genus Ochodaeus Serville with descriptions of two new species in the O. pectoralis LeConte species complex (Coleoptera: Scarabaeidae). Bulletin of the Southern California Academy of Sciences 74: 49-65.

Carlson, D. C., and P. O. Ritcher. 1974. A new genus of Ochodaeinae and a description of the larva of Pseudochodaeus estriatus (Schaeffer). Pan-Pacific Entomologist 50: 99-110.

Eschscholtz, J. F. 1821. Zusätze und Berichtigungen zu den im VI. Bande der Memoires de l’Acad. de Sc. à St. Petersbourg, von mir beschriebenen Insekten. Magazin der Entomologie 4: 397-402.

Fall, H. F. and T.D.A. Cockerell. 1907. The Coleoptera of New Mexico. Transactions of the American Entomological Society 33: 145-272.

Fall, H. F. 1909. A short synopsis of the species of Ochodaeus inhabiting the United States. Journal of the New York Entomological Society 17: 30-38.

Horn, G. H. 1876. Revision of the United States species of Ochodaeus and other genera of Scarabaeidae. Transactions of the American Entomological Society 5: 177-198.

Horn, G. H. 1895. Coleoptera of Baja California, Supplement I. Proceedings of the California Academy of Science 5: 225-259.

Howden, H. F. 1968. Canadian Ochodaeus, with a description of a new species (Coleoptera: Scarabaeidae). Canadian Entomologist 100: 1118-1120.

LeConte, J. L. 1854. Description of the Coleoptera collected by Thomas H. Webb, M.D., in the years. 1850-51 and 52, while Secretary of the U.S. and Mexican Boundary Commission. Proceedings of the Academy of Natural Sciences of Philadelphia 7: 220-225.

LeConte, J. L. 1863. New species of North American Coleoptera. Part 1. Smithsonian Miscellaneous Collections 6(167): 1-92.

LeConte, J. L. 1868. New Coleoptera collected on the survey for the extension of the Union Pacific railway, E.D. from Kansas to Fort Craig, New Mexico. Transactions of the American Entomological Society 2: 49-64.

Linell, M. L. 1896. New species of North American Coleoptera of the family Scarabaeidae. Proceedings of the United States National Museum (1895) 18: 721-731.

Mulsant, E., and C. Rey. 1871. Tribu des Lamellicornes. Annales de la Société d’Agriculture, Sciences et Industries de Lyon, (4)3: 155-480.

Nikolajev, G. V. 1995. Materialy k sistematike podsemeistva Ochodaeinae (Coleoptera, Scarabaeidae). Zoologicheskiy Zhurnal 74: 72-82.

Paulsen, M.J. 2007. Nomenclatural changes in the Nearctic Ochodaeinae and description of two new genera (Coleoptera: Scarabaeoidea: Ochodaeidae). Insecta Mundi 0021: 1-13. pdf DOWNLOAD

Say, T. 1835. Descriptions of new coleopterous insects, and observations on some already described. Boston Journal of Natural History 1: 151-203.

Schaeffer, C. 1906. Four Ochodaeus new to the United States. Canadian Entomologist 38: 269-272.

Scholtz, C. H., D. D’Hotman, A. V. Evans, and A. Nel. 1988. Phylogeny and systematics of the Ochodaeidae (Insecta: Coleoptera: Scarabaeoidea). Journal of the Entomological Society of South Africa 51: 207-240.

Westwood, J. O. 1852. On the lamellicorn beetles which possess exserted mandibles and labrum, and 10-jointed antennae. Transactions of the Entomological Society of London (ns) 2: 59-74.


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Author: M.J. Paulsen - mpaulsen@unlserve.unl.edu
Generated on: 01/JUL/2005
.....Last modified: 07/FEB/2008
UNL State Museum - Division of Entomology