Type genus: Hybosorus MacLeay, 1819.

Description. Length 2.0-15.0 mm. Shape variable, generally rounded. Color brown, black, or reddish brown. Head not deflexed. Antennae 10-segmented (rarely 9-segmented), with 3-segmented, opposable club; club oval or elongate, first segment cupuliform or not. Eye canthus developed. Eyes developed. Labrum produced beyond apex of clypeus, prominent. Mandibles produced beyond apex of labrum, prominent, external edge rounded at base, apex pointed, external surface generally not sculptured. Maxillae with 4-segmented palpi, labium with apex not indented, with 4-segmented palpi. Pronotum convex, generally punctate, base sinuous or rounded. Scutellum exposed, triangular. Elytra convex, generally punctate, striae well-defined. Pygidium concealed by elytra. Hind wings well-developed or reduced, with M-Cu loop present, MP3 and MP4 veins present. Venter with surface generally smooth, sometimes vermiculate. Prosternum bi-concave. Mesosternum invaginated. Abdomen with 5-6 free sternites and 8 functional spiracles, spiracles 1-7 situated on pleural membrane, spiracle 8 situated in tergite. Protibiae with 2 or 3 teeth. Meso- and metatibia with or without transversal carinae. Tarsi 5-5-5. Ventral position of protarsi at or apical to the second tibial tooth. Tarsal claws simple or complex (toothed or with ventral split). Male genitalia asymmetrical.

Composition. The subfamily Hybosorinae includes 24 genera (four fossil) and 132 species (five fossil).

Distribution. Australian, Afrotropical, Indomalaysian, Neotropical, Nearctic, and Palaearctic regions.

Natural history. Adults feed primarily on dung and carrion, and one species is known to be predaceous (Hybosorus illigeri) (Rozas et al. 1991; Ocampo 2002). Most species are attracted to lights. Larvae have been collected in decomposed plant material and associated with roots. Larvae of some species stridulate.

Bibliography:

Ocampo, F. C. 2006. Phylogenetic Analysis of the Scarab Family Hybosoridae and Monographic Revision of the New World Subfamily Anaidinae (Coleoptera: Scarabaeidae). Bulletin of the University of Nebraska State Museum, Vol. 19. 210pp.