HYBOSORIDAE
Erichson, 1847
Anaidinae
Nikolajev, 1996
Cryptogeniinae
Howden, 2001 (new synonym).
Type
genus Anaides Westwood, 1845.
Anaides
Westwood, 1845 (13).
Aneides
(Incorrect subsequent spelling by Balthasar 1838).
Anaides
carioca Ocampo Brazil.
*Anaides
fossulatus Westwood 1846. Panama, Colombia, Venezuela, Guyana, Trinidad
and Tobago, Suriname.
Anaides
reticulatus, Endrödi, 1962. (synonym).
†Anaides
howdeni Ocampo, Dominican Republic.
Anaides laticollis
Harold, 1863. Belize, Costa Rica, El Salvador, Guatemala, Honduras,
Mexico, Nicaragua, Panama.
Anaides longeciliatus Balthasar, 1938. Costa
Rica, Panama.
Anaides
onofrii Ocampo, Colombia, Ecuador, Peru, Bolivia.
Anaides
ortii Ocampo, Venezuela.
Anaides
parvulus Ocampo, Venezuela.
Anaides
planus Ocampo, Costa Rica, Ecuador.
Anaides
quinckei Ocampo Bolivia.
Anaides rugosus Robinson, 1948a. Ecuador,
Peru.
Anaides simplicicollis Bates, 1887. Costa
Rica, Panama.
Anaides
vartorellii Ocampo Barbados.
Callosides
Howden, 1971 (3).
*Callosides
campbelli Howden, 1971. Colombia.
Callosides
bartolozzii Paulian and Cambefort, 1995. Ecuador.
Callosides
genieri Howden, 2001. Ecuador.
Chaetodus
Westwood, 1845 (33).
Subgenus
Chaetodus Westwood, 1845.
Pseudohybosorus
Endrödi, 1963. (synonym).
Chaetodus
allsoppi Martínez, 1988. Peru.
Chaetodus
amazonicus de Borre, 1886. Brazil, Suriname, Venezuela.
Pseudohybosorus drifti Endrödi, 1963 (synonym).
Chaetodus
amazonicus inesperatus Martínez, 1988 (synonym).
Chaetodus
bolivianus Martínez, 1956. Bolivia.
Chaetodus
columbicus Petrovitz, 1970. Colombia.
Chaetodus
datoi Ocampo, Bolivia
Chaetodus
exaratus Arrow, 1909. Brazil, Argentina, Brazil, Paraguay.
Chaetodus apicipennis
Petrovitz, 1970. (synonym).
Chaetodus
fraternus Martínez, 1994. Bolivia.
Chaetodus
globosus Ocampo, Mexico.
Chaetodus
humerosus Petrovitz, 1970. Venezuela.
Chaetodus
hoffmanni Ocampo, Panama, Venezuela.
Chaetodus
irregularis Westwood, 1846. Bolivia, Brazil, Paraguay, Argentina,
Uruguay.
Chaetodus striatus
de Borre, 1886b (synonym).
Chaetodus
jamesonae Ocampo, Venezuela.
Chaetodus
lacandonicus Martínez and Morn, 1990. Mexico, Guatemala.
Chaetodus
maquipucuna Ocampo, Ecuador.
Chaetodus
mimi Ocampo, Ecuador, Peru, Bolivia, Brazil.
Chaetodus
nigrifrons Ocampo, Ecuador.
Chaetodus
noirregularis Ocampo, Brazil.
Chaetodus
paucarae Ocampo, Ecuador.
Chaetodus
paulseni Ocampo, Brazil.
Chaetodus
pax Ocampo, Venezuela.
*Chaetodus
piceus Westwood, 1846. Argentina, Bolivia, Brazil, Paraguay.
Chaetodus
platynotus Ocampo, Ecuador.
Chaetodus
ratcliffei Ocampo, Brazil.
Chaetodus
rodolfo Ocampo, Ecuador.
Chaetodus
sagittarius Ocampo, Brazil.
Chaetodus
smithi Ocampo, Peru, Bolivia, Brazil.
Chaetodus
teamscaraborum Ocampo Belize, Colombia, Costa Rica,
Guatemala, Honduras, Nicaragua, Panama, Venezuela.
Chaetodus
villosicollis Benderitter, 1923. Argentina, Paraguay.
Subgenus
Chaetodopsis Martínez, 1988.
*Chaetodus
asuai Martínez, 1956. Colombia, Ecuador, Peru, Bolivia.
Chaetodus brancuccii Martínez, 1994. Venezuela.
Chaetodus octocarinatus Ocampo, Brazil, Guyana, Suriname.
Chaetodus tricarinatus Ocampo, Colombia, Ecuador, Peru.
Chaetodus venezolanus Martínez, 1994. Venezuela.
Cretanaides
Nikolajev, 1996 (1).
*†Cretanaides
trogopterus Nikolajev, 1996. Siberia (Russia).
Cryptogenius
Westwood, 1845 (2).
Cremastochilodius
Krikken, 1975. (synonym, type species Cremastochilodius trisits
Krikken 1975).
*Cryptogenius
miersianus Westwood, 1846. Costa Rica, Panama, Colombia, Ecuador.
Cryptogenius
fryi Arrow, 1909. Brazil, Argentina.
Cremastochilodius tristis Krikken, 1975. (synonym).
Hybochaetodus Arrow, 1909 (2).
Hybochaetodus
flaco Ocampo, 2002. Peru.
*Hybochaetodus
obscurus Arrow, 1909. Peru.
Totoia
Ocampo, 2003 (3).
Totoia
brachycarina Ocampo, 2003. Nicaragua, Costa Rica. Panama.
*Totoia
splendida Ocampo, 2003. Panama, Colombia.
Totoia
magnifica Ocampo, Panama.
Ceratocanthinae Martínez, 1968.
Acanthocerinae Lacordaire, 1856 (synonym, see below under
Ceratocanthus).
Tribe Ceratocanthini
Martínez, 1968.
Acanthocerodes Péringuey,
1901 (3).
Acanthocerodes endroedyi
Paulian, 1980. South Africa.
Acanthocerodes martini Paulian,
1992. South Africa.
*Acanthocerodes singularis Péringuey, 1901. South Africa.
Afrocloetus
Petrovitz, 1968 (1).
Afrocloeotus (incorrect
subsequent spelling by Paulian, 1977a)
*Afrocloetus gibbosus
Petrovitz, 1968. Tanzania.
Aneilobolus Hesse,
1948 (4).
Aneilobolus endroedyyoungai Paulian, 1983. South
Africa.
Aneilobolus gigas
Paulian, 1977a. South Africa.
*Aneilobolus lawrencei
Hesse, 1948. South Africa.
Aneilobolus leleupi
Paulian, 1977. South Africa.
Anopsiostes
Paulian, 1982 (1)
*Anopsiostes punctatus Paulian, 1982. Ecuador, Peru.
Astaenomoechus
Martínez and Pereira, 1959 (22).
Belloanopsiostes Vaz de Mello, 1996. (synonym).
(type species: B. gilli Vaz de Mello, 1996).
Astaenomoechus americanus (Boucomont, 1936). (Philharmostes).
Colombia, Costa Rica, Ecuador, French Guiana, Mexico, Peru, Trinidad.
Astaenomoechus termitophilus Petrovitz,
1976. (synonym).
Astaenomoechus
andersoni Howden and Gill, 2000. Venezuela.
Astaenomoechus
cavei Howden and Gill, 2003. Belize, Honduras.
Astaenomoechus
cicheroi Martínez, 1969. Bolivia.
Astaenomoechus criberrimus Paulian,
1982. Peru.
Astaenomoechus
estriatus Paulian, 1982. Ecuador.
Astaenomoechus gilli (Vaz de Mello,
1996). (Belloanopsiostes). Brazil.
Astaenomoechus
hispidus Howden and Gill, 2003. Costa Rica.
*Astaenomoechus
hospes (Wasmann, 1902). (Sphaeromorphus). Argentina, Brazil.
Astaenomoechus luniferus Petrovitz,
1973. Colombia, Ecuador.
Astaenomoechus
mixtus Howden and Gill, 2003. Costa Rica, Panama.
Astaenomoechus
multipunctatus Howden and Gill, 2003. Nicaragua,
Costa Rica, Panama.
Astaenomoechus
nevermanni (Boucomont, 1936). (Philharmostes). Costa Rica.
Astaenomoechus
redtenbacheri (Harold, 1874). (Acanthocerus). Colombia,
Venezuela.
Astaenomoechus
paniculus Howden and Gill, 2003. Costa
Rica, Panama.
Astaenomoechus
parvosetosus Howden and Gill, 2003. Costa
Rica, Panama.
Astaenomoechus
punctifrons Howden and Gill, 2003. Nicaragua,
Costa Rica.
Astaenomoechus setosus (Boucomont,
1936). (Philharmostes).
Brazil, Colombia, Costa Rica, Ecuador,
Guyana.
Astaenomoechus
setulosus (Harold, 1874). (Acanthocerus). Colombia, French Guiana,
Venezuela.
Astaenomoechus solisi Howden and Gill, 2003. Nicaragua,
Costa Rica.
Astaenomoechus
strigulosus Howden and Gill, 2003. Costa
Rica
Astaenomoechus unidentatus Petrovitz,
1973. Colombia, Ecuador.
Aulisostes
Howden and Gill, 2000 (2).
Aulisostes paradoxus (Paulian, 1982). (Germarostes). Brazil.
*Aulisostes
pseudoparadoxus Howden and Gill, 2001. Colombia.
Baloghianestes Paulian, 1968 (1).
*Baloghianestes lissoubai
Paulian, 1968. Cameroon, Congo.
Besuchetostes Paulian, 1972 (11).
Besuchetostes besucheti (Paulian, 1972). (Cyphopisthes).
Sri Lanka.
Philharmostes carinatus
Paulian, 1975. (synonym).
Besuchetostes dubium Paulian, 1975.
India.
Besuchetostes hindu Paulian, 1975.
India.
Besuchetostes howdeni Paulian, 1979.
Malaysia.
Besuchetostes jaccoudi Paulian, 1977.
Malaysia.
Besuchetostes keralae Paulian, 1975.
India.
Besuchetostes kodaikanalense Paulian,
1975. India.
Besuchetostes loebli Paulian, 1972.
Sri Lanka.
Besuchetostes mussardi Paulian, 1972.
Sri Lanka.
Besuchetostes peradeniyae Paulian,
1972. Sri Lanka.
*Besuchetostes taprobanae Paulian,
1972. Sri Lanka.
Callophilharmostes
Paulian, 1968 (1).
Philharmostes
(Callophilharmostes) Paulian, 1968
*Callophilharmostes fleutiauxii (Paulian, 1943). (Philharmostes). Cameroon, Gabon, Guinea,
Ivory Coast.
Carinophilharmostes Paulian, 1968 (1).
Philharmostes (Carinophilharmostes)
Paulian, 1968.
*Carinophilharmostes vadoni (Paulian, 1937). (Philharmostes).
Cameroon, Central African
Republic, Congo, Zaire, Equatorial Guinea, Guinea, Ivory Coast,
Uganda.
Pterorthochaetes multigibber Petrovitz, 1967. (synonym).
Ceratocanthoides Paulian, 1982 (1).
*Ceratocanthoides undatus
(Petrovitz, 1973). (Acanthocerus).
Brazil, Colombia, Peru.
Ceratocanthopsis Paulian, 1982 (3).
*Ceratocanthopsis fulgida (Martínez,
1967). (Acanthocerus). Brazil.
Ceratocanthopsis pernitida Paulian,
1982. Bolivia, Brazil, French Guiana, Trinidad.
Ceratocanthopsis pygmaea
(Harold, 1874). (Acanthocerus). Brazil, French Guiana, Venezuela,
Suriname.
Ceratocanthus
White, 1842 (53).
Acanthocerus
MacLeay, 1819. (synonym). (junior homonym of Acanthocerus Palisot
de Beauvois, 1818, Hemiptera)
Gymnoropterus
Gestro, 1899. (synonym). (type species Synarmostes
striatulus Lansberge, 1887).
Sphaeromorphus
Germar, 1843. (synonym).
*Ceratocanthus
aeneus (MacLeay, 1819). (Acanthocerus). USA.
Sphaeromorphus
volvox Erichson in Germar, 1843 (synonym).
Ceratocanthus
amazonicus Paulian, 1982. Brazil, Colombia, French
Guiana.
Ceratocanthus
aureolus (Harold, 1874). (Acanthocerus). Brazil.
Ceratocanthus
baniensis Howden, 1978. Hispaniola.
Ceratocanthus
basilicus (Germar, 1843). (Sphaeromorphus). Brazil.
Ceratocanthus
bicinctoides Paulian, 1982. Brazil.
Ceratocanthus
bicinctus (Germar, 1843). (Sphaeromorphus). Brazil.
Ceratocanthus
bonfilsi Chalumeau, 1977. Guadeloupe.
Ceratocanthus
brasiliensis (Lansberge, 1887). (Acanthocerus). Brazil.
Ceratocanthus
chalceus (Germar, 1843). (Sphaeromorphus). Cuba.
Ceratocanthus
clypealis (Lansberge, 1887). (Acanthocerus). Brazil, French
Guiana, Peru.
Ceratocanthus
ebeninus (Erichson in Germar, 1843). (Sphaeromorphus). Brazil.
Ceratocanthus eberti Paulian, 1982. Brazil.
Ceratocanthus eulampros (Bates, 1887). (Acanthocerus).
Costa Rica, Nicaragua, Panama.
Ceratocanthus
fuscoviridis (Ohaus, 1911). (Acanthocerus). Argentina.
Ceratocanthus globulus (Erichson in Germar, 1843).
(Sphaeromorphus). Brazil, Colombia, French Guiana.
Ceratocanthus
gundlachi (Harold, 1874). (Acanthocerus). Cuba.
Ceratocanthus
humeralis (Erichson in Germar, 1843)). (Sphaeromorphus). Brazil, Panama, Suriname.
Ceratocanthus
inca Paulian, 1982. Peru.
Ceratocanthus
major Paulian, 1982. French Guiana.
Ceratocanthus
mathani Paulian, 1982. Peru.
Ceratocanthus
micans (Harold, 1874). (Acanthocerus). Brazil.
Ceratocanthus
micros (Bates, 1887). (Acanthocerus).
Costa Rica, Mexico.
Ceratocanthus monrosi (Martínez and Pereira,
1959). Bolivia, Brazil.
Ceratocanthus
nanus (Germar, 1843). (Sphaeromorphus). Brazil, Paraguay.
Ceratocanthus niger Paulian, 1982b. Brazil.
Ceratocanthus nitidus (Germar, 1843). (Sphaeromorphus).
Bolivia, Brazil.
Ceratocanthus
pararelucens Howden, 1978. St. Vincent.
Ceratocanthus
pauliani (Delgado and Hernandez, 1998). (Anopsiostes). Mexico.
Ceratocanthus pecki Paulian, 1982. Ecuador.
Ceratocanthus
perpunctatus Paulian, 1982. Ecuador, Peru.
Ceratocanthus politus (Erichson in Germar, 1843).
(Sphaeromorphus). Argentina, Bolivia, Brazil, Peru.
Ceratocanthus pseudosuturalis Paulian, 1982. Brazil.
Ceratocanthus punctolineatus Paulian,
1982. Brazil, Colombia.
Ceratocanthus
punctulatus (Lansberge, 1887). (Acanthocerus). Brazil, Peru.
Ceratocanthus
pyritosus (Erichson in Germar, 1843). (Sphaeromorphus). St.
Thomas.
Ceratocanthus
quadristriatus Paulian and Vaz de Mello, 1998. Brazil.
Ceratocanthus
relucens mexicanus (Bates, 1887). (Acanthocerus). Belize, Mexico.
Ceratocanthus
relucens relucens (Bates, 1887). (Acanthocerus). Belize,
Costa Rica, Guatemala, Mexico, Nicaragua,
Panama.
Ceratocanthus
rotundicollis (Bates, 1887). (Acanthocerus). Panama.
Ceratocanthus
semipunctatus carioca (Martínez, 1967). (Acanthocerus).
Brazil.
Ceratocanthus
semipunctatus semipunctatus (Germar, 1843). (Sphaeromorphus). Brazil,
Colombia, Paraguay.
Ceratocanthus
semistriatus (Germar, 1843). (Sphaeromorphus). Cuba.
Ceratocanthus
seriatus (Erichson in Germar, 1843). (Sphaeromorphus). Brazil,
French Guiana, Peru.
Ceratocanthus
sesquistriatus (Germar, 1843). (Sphaeromorphus). Brazil.
Ceratocanthus
sexstriatus Paulian, 1982. Brazil.
Ceratocanthus spinicornis (Fabricius, 1792). (Trox). Unknown distribution.
Ceratocanthus steinbachi Paulian, 1982b. Bolivia.
Ceratocanthus
striatulus (Lansberge, 1887). (Synarmostes). Unknown distribution.
Ceratocanthus
suturalis (Lansberge, 1887). (Acanthocerus). Bolivia, Brazil,
Ecuador, French Guiana, Peru, Surinam, Trinidad.
Ceratocanthus
suturaloides Paulian, 1982. Venezuela.
Ceratocanthus termiticola (Wasmann, 1894). (Acanthocerus).
Bolivia, Brazil, Colombia, Ecuador.
Ceratocanthus
turquinensis (Zayas, 1988). (Acanthocerus). Cuba.
Ceratocanthus
undulatus (Harold, 1874). (Acanthocerus). Brazil, French
Guiana.
Ceratocanthus
vicarius (Bates, 1887). (Acanthocerus). Belize, Guatemala,
Mexico.
Chaetophilharmostes
Paulian, 1977 (1).
*Chaetophilharmostes chevalieri (Paulian, 1937). (Philharmostes). Congo, Guinea, Ivory Coast,
Liberia.
Pterorthochaetes
termitophilus Petrovitz, 1968. (synonym).
Cloeotus
Germar, 1843 (3).
*Cloeotus
latebrosus Germar, 1843. Colombia.
Cloeotus petrovitzi Paulian, 1982. Brazil.
Cloeotus semicostatus Germar, 1843. Colombia.
Congomostes Paulian, 1968 (2).
*Congomostes baloghi Paulian, 1968. Congo (=former
Zaire.).
Congomostes janssensi (Basilewsky, 1955). (Philharmostes).
Congo (=former Zaire).
Cryptophilharmostes Ballerio, 2000 (2).
*Cryptophilharmostes mahunkai Ballerio, 2000. Tanzania.
Cryptophilharmostes
merkli Ballerio, 2005. Tanzania
Cyphopisthes Gestro, 1899 (10).
*Cyphopisthes
amphicyllis (Sharp, 1875). (Synarmostes). Indonesia, "New
Guinea."
Cyphopisthes
crux (Sharp, 1875). (Synarmostes). "Borneo."
Cyphopisthes
descarpentriesi Paulian, 1977c. Australia.
Cyphopisthes
dohertyi Paulian, 1942. (Philharmostes). India.
Cyphopisthes
inexpectatus Paulian, 1981. Philippines.
Cyphopisthes
krikkeni Paulian, 1980. Indonesia.
Cyphopisthes
luzonicus Paulian, 1978. Philippines.
Cyphopisthes
minutus Paulian, 1978. Malaysia.
Cyphopisthes
szentivanyi (Paulian, 1973). (Philharmostes). Papua New Guinea.
Cyphopisthes
wallacei (Pascoe, 1860). (Sphaeromorphus). Malaysia, Indonesia.
Cyphopisthes humeralis Gestro, 1899 (synonym).
Ebbrittoniella
Martínez, 1962 (2).
Eubrittoniella (incorrect subsequent spelling by Paulian, 1978)
Ebbrittoniella gestroi (Paulian, 1942). (Philharmostes).
Indonesia, Malaysia,
*Ebbrittoniella ignita (Westwood, 1883). (Acanthocerus (Sphaeromorphus). Indonesia,
Malaysia.
Eusphaeropeltis Gestro, 1899 (16).
Eusphaeropeltis aureola Gestro, 1899. Indonesia (Sumatra).
*Eusphaeropeltis aurora (Lansberge, 1887). (Synarmostes).
Indonesia (Sumatra).
Eusphaeropeltis
borneensis Paulian, 1978. "Borneo."
Eusphaeropeltis boucomonti Paulian, 1978. Indonesia (Sumatra).
Eusphaeropeltis celebicus Paulian, 1978. Indonesia (Sulawesi).
Eusphaeropeltis corruscus Gestro, 1899. Malaysia.
Eusphaeropeltis drescheri Paulian, 1942. Indonesia (Java).
Eusphaeropeltis iris Gestro, 1899. Indonesia (Sumatra).
Eusphaeropeltis kedahensis Paulian, 1942. Indonesia (Sumatra).
Eusphaeropeltis krikkeni Paulian, 1978. Indonesia (Sumatra).
Eusphaeropeltis modiglianii Gestro, 1899. Indonesia (Sumatra).
Eusphaeropeltis pulcher (Lansberge, 1885). (Synarmostes).
Indonesia (Sumatra).
Eusphaeropeltis punctatissimus (Lansberge, 1887).
(Synarmostes). Brunei, Malaysia.
Eusphaeropeltis raapi Gestro, 1899. Indonesia (Nias).
Eusphaeropeltis sabah Paulian, 1989. Malaysia (Sabah).
Eusphaeropeltis scheuerni Paulian, 1982a. Indonesia
(Sumatra).
Germarostes Paulian, 1982 (69).
Subgenus Germarostes Paulian, 1982.
Germarostes (Germarostes) allorgei (Paulian, 1947). (Cloeotus).
Dominica, Guadeloupe.
Germarostes (Germarostes) anchicayae Paulian, 1982. Colombia.
Germarostes (Germarostes) antiquus (Erichson in Germar,
1843). (Acanthocerus). Peru, Surinam, Venezuela.
*Germarostes (Germarostes) aphodioides aphodioides (Illiger, 1800). (Melolontha). Canada
to Argentina.
Acanthocerus laevistriatus Laporte, 1840. (synonym).
Scarabaeus latipes Germar, 1824. (synonym).
Trox
splendidus Say, 1835. (synonym).
Germarostes (Germarostes) aphodioides prionomus (Bates, 1887). (Cloeotus).
Guatemala.
Germarostes (Germarostes) argentinus (Ohaus, 1911). (Cloeotus).
Argentina.
Germarostes (Germarostes) bidens (Bates, 1887). (Cloeotus).
Brazil, Panama.
Germarostes (Germarostes) carinatus Paulian, 1982. Ecuador.
Germarostes (Germarostes) carltoni Howden & Gill,
2005. Peru.
Germarostes (Germarostes) costulatus Paulian, 1982. Colombia.
Germarostes (Germarostes) crassicollis (Arrow, 1903). (Cloeotus). St. Vincent.
Germarostes (Germarostes) degallieri Paulian, 1982. Colombia,
Ecuador, French Guiana.
Germarostes (Germarostes) ecuadoricus Paulian, 1982. Ecuador.
Germarostes (Germarostes) excisus (Bates, 1887). (Cloeotus).
Guatemala, Mexico.
Germarostes (Germarostes) farri (Howden, 1970). (Cloeotus). Jamaica.
Germarostes (Germarostes) gaujoni Paulian, 1982. Ecuador.
Germarostes (Germarostes) globosus (Say, 1835). (Trox).
Mexico, USA.
Germarostes (Germarostes) heterodynamus Paulian, 1982. Brazil.
Germarostes (Germarostes) howdenicus Paulian, 1982. Ecuador.
Germarostes (Germarostes) infantulus (Bates, 1887). (Cloeotus).
Guatemala.
Germarostes (Germarostes) instriatus Paulian, 1982. Ecuador.
Germarostes (Germarostes) jamaicensis (Howden, 1978). (Cloeotus).
Jamaica.
Germarostes (Germarostes) leticiae Paulian, 1982.. Brazil, Colombia.
Germarostes (Germarostes) macleayi (Perty, 1830). (Acanthocerus).
Guatemala, Panama, Colombia, French Guiana, Peru, Brazil, Bolivia, Argentina,
Paraguay.
Germarostes (Germarostes) oberthueri Paulian, 1982. Brazil.
Germarostes
(Germarostes) pauliani (Chalumeau
and Cambefort, 1976). (Cloeotus). Guadaloupe
Germarostes
(Germarostes) pecki (Howden, 1970). (Cloeotus). Jamaica.
Germarostes
(Germarostes) plicatus (Erichson in Germar, 1843)). (Acanthocerus).
Mexico to
Argentina except Chile.
Germarostes
(Germarostes) posticus (Germar, 1843). (Acanthocerus). Argentina?,
Chile,
Acanthocerus
muricatus Curtis, 1845. (synonym).
Cloeotus posticus mochae Gutiérrez, 1946 (synonym?).
Germarostes
(Germarostes) pullus Paulian, 1982b. Ecuador
Germarostes
(Germarostes) puncticollis (Erichson in Germar, 1843). (Acanthocerus). Brazil,
Ecuador, Paraguay.
Germarostes
(Germarostes) punctulatus (Ohaus, 1911). (Cloeotus). Argentina, Bolivia, Brazil,
Colombia, French Guiana, Paraguay.
Germarostes (Germarostes) pusillus ( Laporte, 1840).
(Acanthocerus). Colombia.
Germarostes (Germarostes) pustulosus (Lansberge, 1887).
(Cloeotus). Colombia.
Germarostes (Germarostes) reticularis (Bates, 1887). (Cloeotus).
Mexico.
Germarostes (Germarostes) rotundatus Paulian, 1982. Venezuela.
Germarostes
(Germarostes) rufopiceus (Arrow, 1903). (Cloeotus). St. Lucia,
St. Vincent.
Germarostes
(Germarostes) rugatus (Germar, 1843). (Acanthocerus). Colombia.
Acanthocerus
rugosus Germar 1843 (synonym).
Germarostes
(Germarostes) salesiacus Paulian, 1982. Brazil.
Germarostes
(Germarostes) semituberculatus (Germar 1843). (Acanthocerus).
Bolivia,
Brazil,
Paraguay, Peru, Venezuela.
Cloeotus acuarius Petrovitz,
1973. (synonym).
Germarostes (Germarostes) sinuatus sejunctus (Bates,
1887). (Cloeotus). Guatemala.
Germarostes (Germarostes) sinuatus sinuatus (Bates,
1887). (Cloeotus). Belize, Guatemala, Mexico.
Germarostes (Germarostes) sticticus (Erichson in Germar,
1843). (Acanthocerus). Brazil, French Guiana, Nicaragua, Suriname.
Germarostes (Germarostes) strigilateris (Bates, 1887). (Cloeotus).
Mexico.
Germarostes (Germarostes) sulcipennis (Harold, 1875). (Cloeotus).
Peru.
Germarostes (Germarostes) viridis (Lansberge, 1887).
(Cloeotus). Mexico.
Subgenus Haroldostes Paulian, 1982b.
Germarostes
(Haroldostes) abruptus (Petrovitz, 1973). (Cloeotus). Brazil.
Germarostes
(Haroldostes) batesi (Harold, 1874a). (Cloeotus). Brazil,
Bolivia.
Germarostes
(Haroldostes) brunnipes (Germar, 1843). (Acanthocerus). Bolivia,
Ecuador, Paraguay.
Germarostes
(Haroldostes) bugabensis (Arrow, 1903). (Cloeotus). Panama.
Germarostes
(Haroldostes) columbianus Paulian, 1982. Colombia.
Germarostes
(Haroldostes) diffundus (Petrovitz, 1976). (Cloeotus). Brazil, Ecuador, Paraguay.
Germarostes (Haroldostes) ecuadoriensis (Petrovitz, 1976).
(Cloeotus). Ecuador, Venezuela.
Germarostes (Haroldostes) geayi Paulian, 1982. Peru.
Germarostes (Haroldostes) guyanensis Paulian, 1982b. French
Guiana, Panama?
Germarostes (Haroldostes) hamiger (Ohaus, 1911). (Cloeotus).
Argentina, Bolivia.
Germarostes
(Haroldostes) haroldi (Arrow, 1911). (Cloeotus). Colombia, Venezuela.
Cloeotus puncticollis Harold, 1874. (synonym)
(homonym of Germarostes (Germarostes) puncticollis Erichson in
Germar, 1843).
Germarostes (Haroldostes) indigaceus (Germar, 1843). (Acanthocerus).
Brazil, Colombia, Ecuador, Venezuela.
Germarostes (Haroldostes) leprieuri (Germar, 1843). (Acanthocerus).
Brazil, Ecuador, French Guiana.
Germarostes
(Haroldostes) madeiranus Paulian, 1982b. Brazil.
Germarostes
(Haroldostes) malkini Paulian, 1982b. Brazil.
Germarostes
(Haroldostes) metallicus (Harold, 1874a). (Cloeotus). Brazil, Colombia, Ecuador, French Guiana.
Germarostes
(Haroldostes) nasutus (Bates, 1887). (Cloeotus). Mexico.
Germarostes
(Haroldostes) nigerrimus (Blanchard, 1841). (Sphaerelytrus).
Bolivia, Colombia, Ecuador.
Cloeotus
grandis Petrovitz, 1973. (synonym).
Germarostes (Haroldostes) nitens (Guérin-Méneville,
1839). (Acanthocerus). Costa Rica, Brazil,
Honduras, Mexico, Panama.
Cloeotus acutipes
Arrow, 1903. (synonym).
*Germarostes
(Haroldostes) rugiceps (Germar, 1843). (Acanthocerus). Brazil,
Panama?, Paraguay.
Germarostes
(Haroldostes) senegalensis (Laporte, 1840). (Acanthocerus).
Brazil, French Guiana.
Acanthocerus striatus
Germar, 1843. (synonym).
Germarostes
(Haroldostes) tibialis (Petrovitz, 1973). (Cloeotus). Brazil.
Germarostes
(Haroldostes) tubericauda (Bates, 1891). (Cloeotus). Colombia, Ecuador.
Germarostes (Haroldostes) viridipennis (Bates, 1887). (Cloeotus).
Costa Rica, Panama.
Germarostes (Haroldostes) viridulus (Bates, 1887). (Cloeotus).
Mexico.
Glyptogermarostes (1)
Glyptopterus Paulian, 1982 (synonym,
homonym of Glyptopterus de Chaudoir,
1838 Carabidae).
*Glyptogermarostes oberthueri (Paulian, 1982). (Glyptopterus). Brazil.
Goudotostes Paulian, 1979 (1).
*Goudotostes scabrosus
( Laporte, 1840). (Acanthocerus). Madagascar.
Macrophilharmostes
Paulian, 1978 (1).
*Macrophilharmostes major (Paulian, 1975a). (Cyphopishtes). Papua New Guinea
Madrasostes Paulian, 1975 (28).
Madrasostes agostii Paulian, 1993. Malaysia.
Madrasostes boucomonti Paulian, 1978. Indonesia,
Malaysia.
Madrasostes burckhardti Paulian, 1989. Malaysia.
Madrasostes clypeale Paulian, 1993. Indonesia (Sumatra).
Madrasostes depressum Paulian, 1992. Indonesia (Sumatra).
Madrasostes feae Gestro, 1899. India, Kampuchea,
Myanmar, Nepal, Thailand.
Madrasostes franzi Paulian, 1978. Thailand.
Madrasostes
granulatum (Paulian, 1975). (Cloeotus). Papua New Guinea.
Madrasostes inaequale Paulian, 1992. Indonesia (Sumatra).
Madrasostes kazumai hisamatsui Ochi, 1990. Japan.
Madrasostes kazumai kazumai Ochi, Johki, and Nakata, 1990.
Japan.
Madrasostes loebli Paulian, 1981b. Papua New Guinea (Bismarck Isl.).
Madrasostes malayanum Paulian, 1979. Malaysia.
Madrasostes masumotoi Paulian, 1987. Thailand.
*Madrasostes
nigrum Paulian, 1975. India.
Madrasostes orousseti Paulian, 1981. Philippines.
Madrasostes parcepunctatum Paulian, 1989. Malaysia.
Madrasostes punctatum Paulian, 1989. Malaysia.
Madrasostes rafflesi Paulian, 1979c. Malaysia.
Madrasostes reticulatum (Lansberge, 1887). (Synarmostes).
Indonesia (Sulawesi), Philippines.
Madrasostes sabah Paulian, 1989. Malaysia (Sabah).
Madrasostes sculpturatum Paulian, 1989. Indonesia,
Malaysia.
Madrasostes simplex Paulian, 1989. Malaysia.
Madrasostes sumatranum Paulian, 1992a. Indonesia
(Sumatra).
Madrasostes tamil (Paulian, 1975b). (Philharmostes). India.
Madrasostes
thai Paulian, 1987. Thailand.
Madrasostes
thoracicum Paulian, 1989. Malaysia.
Madrasostes
tonkinense (Paulian, 1945). (Cloeotus). Vietnam.
Madrasostes
variolosum (Harold, 1874b). (Cloeotus). Indonesia, Malaysia.
Martinezostes Paulian, 1982 (3).
*Martinezostes asper (F. Philippi, 1859). (Acanthocerus).
Chile.
Martinezostes fortecostatus (Gutiérrez, 1949).
(Cloeotus). Chile.
Martinezostes ruizi (Gutiérrez, 1946).
(Cloeotus). Chile.
Melanophilharmostes Paulian, 1968 (17).
Philharmostes (Melanophilharmostes) Paulian, 1968
Melanophilharmostes ashantii (Paulian, 1974). (Philharmostes).
Ghana.
Melanophilharmostes bicarinatus (Paulian, 1974). (Philharmostes).
Ghana.
Melanophilharmostes burgeoni (Paulian, 1946). (Pterorthochaetes).
Cameroon, Zaire, Angola?
Melanophilharmostes carinatus (Paulian, 1974). (Philharmostes).
Ghana.
Melanophilharmostes carvalhoi (Martínez, 1970). (Astaenomoechus).
Angola.
Melanophilharmostes demirei Paulian, 1977. Cameroon.
Melanophilharmostes donisi (Basilewsky, 1955). (Pterorthochaetes).
Zaire.
Melanophilharmostes endroedyi (Paulian, 1968. (Philharmostes
(Melanophilharmostes) Zaire.
Melanophilharmostes ghanae (Paulian, 1974). (Philharmostes).
Ghana, Togo.
Melanophilharmostes
ocellatus (Paulian, 1968). (Philharmostes (Melanophilharmostes). Congo.
Melanophilharmostes palustris (Petrovitz, 1968). (Philharmostes).
Zaire.
Melanophilharmostes puncticeps (Paulian, 1946). (Pterorthochaetes).
Zaire, Liberia.
Melanophilharmostes posthi (Paulian, 1937). (Pterorthochaetes).
Ivory Coast, Liberia, Togo.
Melanophilharmostes pseudoposthi (Paulian, 1977). Zaire.
Melanophilharmostes pygmaeus (Petrovitz, 1968). (Philharmostes).
Zaire.
Melanophilharmostes vincenti (Paulian, 1968). (Philharmostes
(Melanophilharmostes). Congo.
*Melanophilharmostes zicsii (Paulian, 1968). (Philharmostes (Melanophilharmostes). Cameroon, Congo, Gabon.
Nesopalla
Paulian and Howden, 1982 (2).
Nesopalla
borinquensis Paulian and Howden, 1982. Puerto
Rico.
*Nesopalla iviei Paulian and Howden, 1982.
Puerto Rico, Virgin Islands.
Paulianostes Ballerio, 2000 (3).
Paulianostes acromialis (Pascoe, 1860). (Sphaeromorphus).
Malaysia.
Philharmostes
arrowi Paulian, 1942 (synonym).
*Paulianostes
georyssoides (Gestro, 1899). (Cyphopisthes). Indonesia, Malaysia.
Paulianostes
panggoling Ballerio, 2000. Brunei, Malaysia (Sabah).
Perignamptus
Harold, 1877 (4 ).
Perignamptus
carinipennis Gestro, 1899. Papua New Guinea.
Perignamptus loriae Gestro, 1899. Papua New Guinea.
Perignamptus rossi Paulian, 1978. Indonesia (Irian
Jaya).
*Perignamptus
sharpi Harold, 1877. Papua New Guinea.
Petrovitzostes Paulian, 1977 (1).
*Petrovitzostes guineensis (Petrovitz, 1968). (Pterorthochaetes). Cameroon, Equatorial
Guinea.
Philharmostes Kolbe, 1895 (31).
Subgenus Philharmostes Kolbe, 1895.
Philharmostes
(Philharmostes) adami Paulian, 1968. (Philharmostes (Carinophilharmostes).
Congo, Guinea, Ivory Coast.
*Philharmostes
(Philharmostes) aeneoviridis Kolbe, 1895. Madagascar.
Philharmostes (Philharmostes)
basicollis (Fairmaire, 1897). Madagascar.
Philharmostes obscurus
Fairmaire, 1900 (synonym).
Philharmostes (Philharmostes) basilevskyi Paulian, 1977. Tanzania.
Philharmostes (Philharmostes) bicolor Boucomont, 1937. Madagascar.
Philharmostes (Philharmostes) boucomonti Petrovitz, 1968. Madagascar.
Philharmostes (Philharmostes) caffer
(Martínez, 1970) (Astaenomoechus). Angola.
Philharmostes (Philharmostes) corruscus Fairmaire, 1903. Madagascar.
Philharmostes pilula Fairmaire, 1903 (synonym).
Philharmostes convexifrons Fairmaire, 1903 (synonym).
Philharmostes (Philharmostes) criberrimus Paulian, 1979. Madagascar.
Philharmostes (Philharmostes) cribrarius Fairmaire, 1903. Madagascar.
Philharmostes (Philharmostes) cupreolus Fairmaire, 1900. Madagascar.
Philharmostes (Philharmostes) descarpentriesi Paulian, 1979. Madagascar.
Philharmostes (Philharmostes) disparilis Hesse, 1948. South Africa.
Philharmostes (Philharmostes)
garciabesi (Martínez, 1970) (Astaenomoechus). Congo
(= former Zaire)
Philharmostes
(Philharmostes) girardi Paulian, 1993. Guinea.
Philharmostes (Philharmostes) grebennikovi Ballerio, 2004. Tanzania
Philharmostes (Philharmostes) integer Kolbe, 1895. Tanzania.
Philharmostes (Philharmostes) interruptus Hesse, 1948. South Africa.
Philharmostes (Philharmostes) latericostatus (Fairmaire, 1885).
(Synarmostes). Madagascar.
Synarmostes obscuroaeneus Fairmaire, 1897 (synonym).
Synarmostes perroti Wasmann, 1897 (synonym).
Philharmostes (Philharmostes) olsoufieffi Boucomont, 1937. Madagascar.
Philharmostes (Philharmostes) ornatus Ballerio, 2004. Tanzania
Philharmostes (Philharmostes) perrieri (Fairmaire, 1898).
(Synarmostes). Madagascar.
Philharmostes (Philharmostes) pseudobasicollis Paulian, 1979. Madagascar.
Philharmostes (Philharmostes) pseudumbratilis Ballerio, 2004. Tanzania
Philharmostes (Philharmostes) spelaeus Paulian, 1979. Zaire.
Philharmostes (Philharmostes) umbilicatus Petrovitz, 1968. Zaire, Guinea
Philharmostes (Philharmostes) umbratilis Petrovitz, 1968. Tanzania.
Philharmostes (Philharmostes) vadonianus Paulian, 1979. Madagascar.
Philharmostes (Philharmostes) werneri Ballerio, 2001. Tanzania.
Philharmostes (Philharmostes) zuluensis Hesse, 1948. South
Africa.
Subgenus Holophilarmostes Paulian, 1968.
*Philharmostes (Holophilharmostes) badius (Petrovitz, 1967).
(Pterorthochaetes). Cameroon,
Congo, Zaire, Guinea.
Philharmostes (Holophilharmostes) cohici Paulian, 1968. (synonym).
Pseudopterorthochaetes Paulian, 1977 (7).
Pseudopterorthochaetes cambeforti Paulian, 1981. Ivory
Coast.
Pseudopterorthochaetes criberrimus Paulian,
1977. Zaire, Gabon.,
*Pseudopterorthochaetes elytratus (Paulian,
1946). (Pterorthochaetes). Cameroon, Zaire.
Pseudopterorthochaetes endroedyi (Paulian,
1974). (Pterorthochaetes). Ghana, Ivory Coast.
Pseudopterorthochaetes hystrix Paulian, 1991.
Madagascar.
Pseudopterorthochaetes kumasii (Paulian, 1974).
(Pterorthochaetes). Gabon, Ghana.
Pseudopterorthochaetes machadoi (Martínez,
1970). (Astaenomoechus). Angola.
Pterorthochaetes Gestro, 1899 (21).
Pterorthochaetes andamanus Paulian, 1937. India (Andaman
Islands).
Pterorthochaetes armatus Paulian, 1945. Vietnam.
Pterorthochaetes
brevis (Sharp, 1875). (Synarmostes). Indonesia (Irian Jaya),
Papua New Guinea.
Acanthocerus
(Sphaeromorphus) byrrhoides Westwood, 1883 (synonym).
Pterorthochaetes
brevisetosus Gestro, 1899. Singapore.
Pterorthochaetes coomani Paulian, 1945. Vietnam.
Pterorthochaetes
cribricollis Gestro, 1899. Australia. Papua New Guinea.
*Pterorthochaetes gestroi gestroi (Harold, 1874).
(Synarmostes).
Indonesia, Malaysia.
Pterorthochaetes gestroi longisetosus Gestro, 1899. Singapore.
Pterorthochaetes
haroldi (Sharp, 1875). (Synarmostes).
Indonesia, Malaysia, Singapore.
Pterorthochaetes hirtus Gestro, 1899. Indonesia (Sumatra).
Pterorthochaetes incertus Gestro, 1899. Malaysia.
Pterorthochaetes
insularis Gestro, 1899. Indonesia, Malaysia, Nepal, Thailand.
Pterorthochaetes
latus (Sharp, 1875). (Synarmostes).
Indonesia (Sumatra), Singapore.
Pterorthochaetes lavongai Paulian, 1968b. Papua New Guinea (Bismarck Isl.).
Pterorthochaetes
laxepunctatus Paulian, 1973. Papua New Guinea.
Pterorthochaetes mallicoloi Paulian, 1968b. Vanuatu Island.
Pterorthochaetes montanus Ballerio, 1999. Malaysia.
Pterorthochaetes picinus (Sharp, 1875). (Synarmostes).
Micronesia, Philippines.
Pterorthochaetes puncticollis (Sharp, 1875). (Synarmostes).
Indonesia (Java).
Pterorthochaetes septemtrionalis Ballerio, 1999. Nepal.
Pterorthochaetes simplex Gestro, 1899. Australia, Papua
New Guinea.
Pterorthochaetes sulawesii Paulian 1987. Indonesia (Sulawesi).
Synarmostes Germar, 1843 (4).
Synarmostes antsingyi Paulian, 1979. Madagascar.
Synarmostes humilis Fairmaire, 1893. Comores islands.
Synarmostes niger Paulian, 1979. Madagascar.
*Synarmostes tibialis (Klug, 1832). (Acanthocerus).
Madagascar.
Tribe Ivieolini Howden and Gill, 2000.
Ivieolus
Howden and Gill, 1988 (3).
Ivieolus
brooksi Howden and Gill, 2000. French Guiana.
Ivieolus
inflaticollis Howden and Gill, 2001. Ecuador.
*Ivieolus
pseudoscutellatus Howden and Gill, 1988. Guyana, Venezuela.
Tribe Scarabatermitini Nikolajev, 1999
Scarabaeinus Silvestri, 1940 (1).
*Scarabaeinus termitophilus Silvestri, 1940. Brazil.
Scarabatermes Howden, 1973 (1).
*Scarabatermes amazonensis Howden, 1973. Colombia.
Trachycrusus
Howden and Gill, 1995 (2).
*Trachycrusus
lescheni Howden and Gill, 1995. Peru.
Trachycrusus
striatulus Howden and Gill, 1995. Peru.
Xenocanthus
Howden and Gill, 1988 (1).
*Xenocanthus singularis
Howden and Gill, 1988. Venezuela.
Hybosorinae
Erichson, 1847.
Type
genus Hybosorus MacLeay, 1819.
Apalonychus
Westwood, 1845 (4).
Hapalonychus
(incorrect subsequent spelling by Lacordaire 1856).
Haplonychus
(incorrect subsequent spelling by Petrovitz (1973).
Trichops
Val, 1853. (synonym). (type sepecies Trichops testaceus Val,
1853)
Apalonychus
nattereri Petrovitz, 1973. Brazil.
Apalonychus
pusillus Arrow, 1911. Paraguay.
Apalonychus
rufulus ( Laporte, 1840). (Hybosorus). Dominican Republic.
*Apalonychus
waterhousei Westwood, 1846. Cuba, Dominican Republic.
Trichops testaceus
Val, 1853. (synonym)
Apalonychus rufulus
de Borre, 1886. (synonym).
Araeotanypus
Waterhouse, 1875 (6).
Araeotanopus
(Incorrect subsequent spelling by Arrow (1912)).
Araeotanypus
bicolor Petrovitz, 1967. South Africa.
*Araeotanypus
boops Waterhouse, 1875. Botswana, South Africa.
Araeotanypus
consors Péringuey, 1908. South Africa.
Araeotanypus
pentheri Petrovitz, 1967. South Africa.
Araeotanypus
striatus Schmidt, 1912. Tanzania.
Araeotanypus
zumpti Petrovitz, 1967. South Africa.
Celaenochrous
Kuijten, 1984 (1).
*Celaenochrous
sinensis Kuijten, 1984. China.
Coilodes
Westwood, 1845 (9).
Coelodes
(incorrect subsequent spelling by Lacordaire (1856))
Coilodes
castaneus Westwood, 1846. Colombia, Costa Rica, Nicaragua.
Coilodes
chilensis Westwood, 1846. Chile? (doubtfull distribution).
Coilodes
fumipennis Arrow, 1909. Brazil.
*Coilodes
gibbus (Perty, 1830). (Hybosorus). Brazil.
Coilodes testaceus
Pic, 1928. (synonym).
Hybosorus brasiliensis
(Laporte, 1840). (synonym).
Coilodes
humeralis (Mannerheim, 1829). (Hybosorus). Brazil.
Hybosorus niger (Mannerheim, 1829). (synonym).
Hybosorus auger Westwood,
1846. (synonym).
Coilodes
nigripennis Arrow, 1903. St. Vincent.
Coilodes
ovalis Robinson, 1948. Venezuela.
Coilodes
parvulus Westwood, 1846. Brazil.
Coilodes
punctipennis Arrow, 1909. Ecuador, Peru.
Coprologus Heer, 1847 (1).
*†Coprologus
gracilis Heer, 1847. Germany.
Cretohybosorus
Nikolajev, 1999 (2).
*†Cretohybosorus
buryaticus Nikolajev, 1999. Russia (Siberia).
†Cretohybosorus
striatulus Nikolajev, 1999. Russia (Siberia).
Dicraeodon
Erichson, 1847 (3).
Aporolaus
Bates, 1887 (synonym).
*Dicraeodon
basalis (Westwood, 1846). (Chaetodus? West.). French Guiana.
Dicraeodon
fimbriatus (Bates, 1887). (Aporolaus). Panama.
Dicraeodon
punctatus Arrow, 1911. Guatemala, Colombia.
Hapalonychoides
Martínez, 1994 (1).
*Hapalonychoides
similis Martínez, 1994. Argentina, Paraguay.
Hybosoroides
Benderitter, 1914 (1).
*Hybosoroides
alluaudi Benderitter, 1914. Kenya.
Hybosorus
MacLeay, 1819 (5).
Hybosorus
crassus Klug, 1855. Congo, Zimbabwe, Mozambique.
*Hybosorus
illigeri Reiche, 1853. Africa, U.S.A., Mexico, Cuba, Haiti, Bahamas,
Nicaragua, Venezuela, Israel, Saudi Arabia, Iran, Central Asia, Afghanistan,
Pakistan, India, China, Vietnam, Portugal, Spain, France, Italy, Hungary,
Greece, Bulgaria, Cyprus.
Scarabaeus arator (Illiger, 1803) homonym
of Scarabaeus arator (Fabricius 1792)
Hybosorus
pinguis Westwood, 1845. (synonym).
Hybosorus
roei Westwood, 1845. (synonym).
Hybosorus
carolinus LeConte, 1847. (synonym).
Hybosorus
nitidus Lansberge, 1882. (synonym).
Hybosorus
illigeri nossibianus Fairmaire, 1895 (synonym).
Hybosorus.
illigeri palearcticus Endrödi, 1957. (synonym).
Hybosorus
laportei Westwood, 1845. Egypt, Ethiopia, Mauritania, Mali, Nigeria,
Senegal, Togo.
Hybosorus thoracicus Westwood, 1845. (synonym)
Hybosorus
orientalis Westwood, 1845. Afghanistan, Myanmar, India, Indonesia
(Java), Pakistan, Sri Lanka, Indonesia.
Hybosorus ruficornis
Boheman, 1857. Botswana, Kenya, Mozambique, Namibia,Somalia, South Africa,
Zambia, Zimbabwe.
Hypseloderus Fairmaire,
1893 (1).
*Hypseloderus
denticollis Fairmaire, 1893a. Vietnam.
Kuijtenous
Paulian, 1981 (2).
*Kuijtenous
laeviceps (Fairmaire, 1893). (Hybosorus). Comores Islands,
Madagascar.
Hybosorus
baliensis Brancsik, 1893 (synonym).
Phaeochrous insularis
Linell, 1897 (synonym)
Kuijtenous
tenuepunctatus (Fairmaire, 1895) (Hybosorus). Madagascar.
Hybosorus
sparsepunctatus Pic, 1930 (synonym).
Metachaetodus de Borre, 1886 (2).
Metachaetodus
brunneicollis de Borre, 1886, Argentina.
*Metachaetodus
discus de Borre, 1886, Uruguay.
Microphaeochroops
Pic, 1930 (5).
Microphaechroops
Pic 1930. (inadvertent error)
*Microphaeochroops
hirsutus Pic, 1930. Vietnam.
Microphaeochroops
laetus Arrow, 1942. Malaysia.
Microphaeochroops
nigrosetosus Kuijten, 1985. Malaysia.
Microphaeochroops
peninsularis Arrow, 1942. Malaysia.
Microphaeochroops
varius Kuijten, 1985. Malaysia.
Microphaeolodes
Kuijten, 1985 (1).
*Microphaeolodes
mulumontis Kuijten, 1985. Malaysia.
Mimocoelodes
Pic, 1930 (1).
*Mimocoelodes
minutus Pic, 1930. Vietnam.
Pantolasius
Lansberge, 1887 (2).
Pantolasius
bandaharae Kuijten, 1985. Indonesia.
*Pantolasius
vethi Lansberge, 1887. Indonesia.
Phaeochridius
Lansberge, 1887 (2).
*Phaeochridius
derasus (Harold, 1880). (Liparochrus).
Indonesia.
Phaeochridius haroldi
Fairmaire, 1896. (synonym)
Phaeochridius
uniformis Arrow, 1925. Malaysia (Sabah).
Phaeochridius benderitteri Pic, 1928. (synonym)
Phaeochridius cinereicollis Arrow, 1942. (synonym)
Phaeochroops
Candze, 1876 (26).
Paeochroops
(incorrect subsequent spelling by Nomura (1973)
Phaechroops
(incorrect subsequent spelling by Pic (1928)
Phaeochroops
acuticollis (Arrow, 1907) (Phaeochridius). Malaysia (Sabah).
Phaeochroops
angulatus Benderitter, 1923. Philippines.
Phaeochroops
colopacilis Kuijten, 1981. Vietnam.
Phaeochroops
curtulus Schmidt, 1912. India.
Phaeochroops
freenae Kuijten, 1981. Malaysia.
Phaeochroops
gigas Arrow, 1907. Malaysia.
Phaeochroops
gilleti Benderitter, 1923. Malaysia, Sabah, Philippines.
Phaeochroops punctulatus
Arrow, 1942. (synonym).
Phaeochroops
hisaeae Nishikawa, 1989. Malaysia, Sabah, Indonesia.
Phaeochroops
indicus Arrow, 1907. India.
Phaeochroops
lakhonicus Kuijten, 1981. Thailand.
*Phaeochroops
lansbergei Candèze, 1876. Indonesia.
Phaeochroops
laotianus Paulian, 1945. Laos.
Phaeochroops
longisetosus Kuijten, 1981. India.
Phaeochroops
maruyamai Nishikawa, 1989. Malaysia, Sabah, Indonesia.
Phaeochroops
masumotoi Nishikawa, 1996. Thailand.
Phaeochroops
meghalayicus Keith, 2001. India.
Phaeochroops
myanmarensis Keith, 2000. Myanmar.
Phaeochroops
ninbin Kuijten, 1981b. Vietnam.
Phaeochroops
opacicollis Arrow, 1909. Myanmar, Thailand.
Phaeochroops
peninsularis Arrow, 1909. Malaysia.
Phaeochroops
rattus Arrow, 1909. Indonesia, Malaysia.
Phaeochroops
batuensis Arrow, 1909. (synonym).
Phaeochroops
mentaweiensis Arrow, 1909. (synonym).
Phaeochroops
niasianus Arrow, 1909. (synonym).
Phaeochroops
seres Kuijten, 1981. Vietnam.
Phaeochroops
silphoides Fairmaire, 1898. Indonesia, Malaysia, Sabah, Philippines.
Phaeochroops recticollis
Pic, 1928. (synonym).
Phaeochroops
taiwanus Nomura, 1973. China (Taiwan).
Phaeochroops
vulpecula Arrow, 1909. Indonesia, Malaysia.
Phaeochroops
vulturius Kuijten, 1981. Indonesia.
Phaeochrous
Laporte, 1840 (51).
Silphodes
Westwood, 1845 (synonym).
Phaeochrous
amplus Arrow, 1909. Cameroon.
Phaeochrous
colmanti Schouteden, 1918 (synonym).
Phaeochrous
bayeri Schouteden, 1918 (synonym).
Phaeochrous
confusus Schouteden, 1918 (synonym).
Phaeochrous
westwoodi Schouteden, 1918 (synonym).
Phaeochrous
australicus Kuijten, 1978. Australia.
Phaeochrous
beccarii Harold, 1871. Ethiopia, Sudan, Tanzania, Yemen.
Phaeochrous
arabicus Arrow, 1909 (synonym).
Phaeochrous
elgonensis Schouteden, 1918 (synonym).
Phaeochrous
gonsalvesi Petrovitz, 1975 (synonym).
Phaeochrous
niloticus Burgeon, 1928 (synonym).
Phaeochrous
stercorarius Kolbe, 1895 (synonym).
Phaeochrous
tangensis Schouteden, 1918 (synonym).
Phaeochrous
becarii, Schmidt, 1913 (synonym).
Phaeochrous
bicarinatus Kuijten, 1986. Uganda, Rwanda, Burundi
Phaeochrous
borealis Kuijten, 1984. China.
Phaeochrous
burgoblitus Kuijten, 1986. Zaire, Uganda, Rwanda.
Phaeochrous kivuensis
Burgeon. (nomen nudum).
Phaeochrous camerunensis
Arrow, 1909. Cameroon, Gabon, Nigeria, Congo, Zaire, Sudan, Ethiopia,
Tanzania, Zambia, Rwanda, Burundi, Uganda, Mozambique, South Africa.
Phaeochrous boranus
Mller, 1939 (synonym).
Phaeochrous
mashunus Arrow 1909 (synonym).
Phaeochrous
zombensis Schouteden, 1918 (synonym).
Phaeochrous vicinus
Schouteden, 1918 (synonym).
Phaeochrous
compactus Kuijten, 1978. Sri Lanka.
Phaeochrous
davaonis Kuijten, 1981. Philippines.
Phaeochrous
dispar Quedenfeldt, 1884. Angola, Zaire.
Phaeochrous thomensis
Arrow, 1909 (synonym).
Phaeochrous kuiluensis
Schouteden, 1918 (synonym).
Phaeochrous
dissimilis dissimilis Arrow, 1909. Myanmar, China, Laos,
Thailand, Vietnam.
Phaeochrous
dissimilis vietnamicola Kuijten, 1978. China, Vietnam.
Phaeochrous
diversipes Pic, 1928. India.
Phaeochrous
dubius (Westwood, 1846). (Silphodes). Indonesia.
Phaeochrous
elevatus Kuijten, 1978. Sri Lanka.
Phaeochrous
emarginatus benderitteri Pic, 1928. Indonesia (Waigeu Island).
Phaeochrous rugosicollis Benderitter, 1913. (synonym).
Phaeochrous celebensis var. ruficeps Pic. 1928. (synonym).
Phaeochrous
emarginatus buruensis Kuijten, 1978. Indonesia (Buru Island)
Phaeochrous
emarginatus davidis Fairmaire, 1886. China.
*Phaeochrous emarginatus
emarginatus Laporte, 1840. Japan, Melanesia, China, India, Thailand,
Buthan, Laos, Vietnam, Malaysia, Indonsesia, Philipines, Papua New Guinea,
Australia.
Phaeochrous sumatrensis
(Westwood, 1846) (Silphodes). (synonym).
Phaeochrous hirtipes (MacLeay,
1864) (Silphodes). (synonym).
Phaeochrous alternatus Fairmaire,
1879 (synonym).
Phaeochrous asiaticus Lewis,
1896 (synonym).
Phaeochrous pallidus Arrow,
1909 (synonym).
Phaeochrous celebensis Pic,
1928 (synonym).
Phaeochrous perroudi Pic,
1943 (synonym).
Phaeochrous gracilis Petrovitz,
1975 (synonym).
Phaeochrous
emarginatus thilliezi Keith, 1999. Pakistan.
Phaeochrous
emarginatus suturalis Lansberge, 1885. (Myanmar).
Phaeochrous
enigmaticus Kuijten, 1978. Bangladesh, China, Indonesia.
Phaeochrous
gambiensis (Westwood, 1846) (Silphodes). Gambia, Senegal,
Burkina, Chad.
Phaeochrous
gigas Schouteden, 1918. Africa.
Phaeochrous latus
Pic, 1944 (synonym).
Phaeochrous
hainanensis Zhang-Youwei, 1990. China.
Phaeochrous
indicus (Westwood, 1846). (Silphodes). India.
Phaeochrous
intermedius intermedius Pic, 1928. Bhutan, China, India,
Laos, Thailand, Vietnam.
Phaeochrous
intermedius occidentalis Kuijten, 1978. Bhutan.
Phaeochrous
lobatus Kuijten, 1978. Philippines.
Phaeochrous
madagascariensis (Westwood, 1846). (Silphodes). Madagascar.
Phaeochrous
madrassicus Kuijten, 1978. India.
Phaeochrous
nicolasi Keith, 2002. Australia.
Phaeochrous
nitidus Arrow, 1909. Tanzania.
Phaeochrous arrowi
Schouteden 1918 (synonym).
Phaeochrous
philippinensis (Westwood, 1846) (Silphodes). Philippines.
Phaeochrous
pletus Kuijten, 1978. East Timor.
Phaeochrous
portuum Kuijten, 1978. New Guinea.
Phaeochrous
pseudintermedius Kuijten, 1978. Vietnam.
Phaeochrous
rhodesianus Schouteden, 1918. Zimbabwe.
Phaeochrous kapiriensis
Schouteden, 1918 (synonym).
Phaeochrous
rudis Kuijten, 1984. Indonesia.
Phaeochrous
ruficollis Fairmaire, 1893a. Vietnam.
Phaeochrous
rufus Pic, 1928b. China.
Phaeochrous
schoutedeni Burgeon, 1928. Central Africa.
Phaeochrous
separabilis Zhang-Youwei, 1990. China.
Phaeochrous
stupendus Kuijten, 1986. Equatorial Africa.
Phaeochrous
sulawesi Kuijten, 1978. Indonesia.
Phaeochrous
tanzanicus Tagliaferri, 2002. Tanzania.
Phaeochrous
tonkineus Pic, 1943. Vietnam.
Phaeochrous tumbanus
Burgeon, 1928. Angola, Ivory Coast, Congo, Zambia, Central African Republic,
Gabon, Ethiopia, Gabon, Kenya, Rwanda, Burundi, Tanzania, Uganda.
Phaeochrous
tokaraensis Nomura, 1961. Japan.
Phaeochrous
turcanicola Kuijten, 1986. Kenya.
Phaeochrous uelensis
Burgeon, 1928. Gabon, Cameroon, Zaire, Uganda, Mozambique, Madagascar.
Phaeochrous
usambarae Burgeon, 1928. Tanzania.
Phaeocroides
Péringuey, 1908 (5).
Phaeochroides
(incorrect subsequent spelling by Schmidt (1913).
*Phaeocroides
damarinus Péringuey, 1908. Namibia.
Phaeocroides
effetus (Kolbe, 1907). (Phaeocrhous?). Namibia, Botswana.
Phaeocroides juvenilis Kolbe, 1912. (synonym).
Phaeocroides
nanus (Arrow, 1942). (Phaeocrhous). India, Pakistan.
Phaeocroides
mapellii Petrovitz, 1972. Bangladesh.
Phaeocroides
orientalis (Petrovitz, 1963). (Araeotanopus).
India.
Procoilodes
Ocampo, 2002 (1).
*†Procoilodes
adrastus Ocampo, 2002. Dominican Republic.
Seleucosorus
Kuijten, 1983 (1).
*Seleucosorus
punctatissumus (Reiche, 1861) (Hybosorus). Syria,
Turkey.
Tyrannasorus
Ratcliffe and Ocampo, 2001 (1).
*†Tyrannasorus
rex Ratcliffe and Ocampo, 2001. Dominican Republic.
Liparochrinae
Ocampo
Type
species Liparochrous Erichson, 1848.
Antiochrus
Sharp, 1873 (7).
Antiochrus
aberrans (Fairmaire, 1877). (Liparaochrus). Australia.
*Antiochrus
brunneus Sharp, 1873. Australia.
Antiochrus oblongus
Harold, 1873 (synonym).
Antiochrus
freyi Petrovitz, 1963. Australia.
Antiochrus
politulus (MacLeay, 1888). (Liparaochrus). Australia.
Antiochrus
septentrionalis Petrovitz, 1968. Australia.
Antiochrus
setosus Petrovitz, 1963. Australia.
Antiochrus
similis Petrovitz, 1971. Australia.
Liparochrus
Erichson, 1848 (41).
Ranidia
Paulian, 1980 (synonym).
Subgenus
uncertain.
Liparochrus
asperulus Fairmaire, 1877. Australia.
Subgenus
Liparochrus Erichson 1848.
Liparochrus
(Liparochrus) bimaculatus Macleay, 1864. Australia.
Liparochrus
(Liparochrus) crassicollis Arrow, 1925. Papua New Guinea.
Liparochrus
(Liparochrus) dilatatifrons Blackburn, 1905. Australia.
Liparochrus dilatifrons
Arrow, 1912. (misspelling).(synonym).
Liparochrus timidus
Arrow, 1909. (synonym).
Liparochrus
(Liparochrus) dolium Heller, 1912. Papua New Guinea.
Liparochrus
(Liparochrus) dux Arrow, 1909. Papua New Guinea.
Liparochrus
(Liparochrus) eungellae Paulian, 1980. Australia.
Liparochrus
(Liparochrus) fossulatus Westwood, 1852. Australia.
*Liparochrus
(Liparochrus) geminatus Westwood, 1852. Australia.
Liparochrus
(Liparochrus) hackeri Blackburn, 1912. Australia.
Liparochrus freyi Petrovitz,
1963. (synonym).
Liparochrus
(Liparochrus) insularis Petrovitz, 1980, Lord Howe Island.
Liparochrus
(Liparochrus) krikkeni Paulian, 1980. Papua New Guinea.
Liparochrus
(Liparochrus) laevipennis Petrovitz, 1963. Australia.
Liparochrus
(Liparochrus) laevis Paulian, 1980. Australia.
Liparochrus
(Liparochrus) laevissimus Paulian, 1980a. Australia.
Liparochrus
(Liparochrus) matthewsi Paulian, 1980a. New Caledonia.
Liparochrus
(Liparochrus) nanus Paulian, 1980a. Australia.
Liparochrus
(Liparochrus) nitidicollis Blackburn, 1905. Australia.
Liparochrus
(Liparochrus) occidentalis Paulian, 1980. Australia.
Liparochrus
(Liparochrus) papuus Lansberge, 1885. Papua New Guinea.
Liparochrus alternatus
Macleay, 1886 (synonym).
Liparochrus
(Liparochrus) quadrimaculatus Harold, 1877. Australia.
Liparochrus
(Liparochrus) rufus Blackburn, 1892. Australia.
Liparochrus
(Liparochrus) sculptilis Westwood, 1852. Australia.
Liparochrus ciliboides
Harold, 1875 (synonym).
Liparochrus
(Liparochrus) septentrionalis Paulian, 1980. Australia.
Liparochrus
(Liparochrus) silphoides Harold, 1874. Australia.
Liparochrus raucus
Fairmaire, 1877 (synonym).
Liparochrus
(Liparochrus) sulcatus (Montrouzier, 1860). (Sphaeridium).
New Caledonia.
Sphaeridium sulcatus Montrouzier, 1860 (synonym).
Subgenus
Parolichrus Allsopp, 1982.
*Liparochrus
(Parolichrus) ingens Felsch, 1909. Papua New Guinea.
Liparochrus ingenti,
Heller 1912. (synonym).
Liparochrus
(Parolichrus) lugubris Arrow, 1925. Indonesia, Papua New Guinea.
Subgenus
Ranidichruss Allsopp, 1982.
Liparochrus
(Ranidichrus) cahilli Paulian, 1980. Australia.
Liparochrus
(Ranidichrus) carnei Paulian, 1980. Australia.
Liparochrus
(Ranidichrus) crenatulus Fairmaire, 1877. Australia.
Liparochrus crenulatus,
Lansberge, 1885. (synonym).
Liparochrus weyersi
Petrovitz, 1971. (synonym).
Liparochrus
(Ranidichrus) darlingtoni Paulian, 1980. Australia.
Liparochrus
(Ranidichrus) demarzi Petrovitz, 1963. Australia.
Liparochrus
(Ranidichrus) globuliformis Macleay, 1888. Australia.
Liparochrus
(Ranidichrus) infantus Petrovitz, 1963. Australia.
Liparochrus
(Ranidichrus) irianae Paulian, 1980. Indonesia, Papua New Guinea.
Liparochrus
(Ranidichrus) modestus Paulian, 1980. Papua New Guinea.
*Liparochrus
(Ranidichrus) multistriatus Harold, 1874. Australia.
Liparochrus pimelioides
Lansberge, 1885. (synonym).
Liparochrus pimeloides,
Schimdt, 1913. (misspelling).
Liparochrus
(Ranidichrus) septemdecimlineatus Petrovitz, 1968. Australia.
Liparochrus
(Ranidichrus) tropicus Petrovitz, 1963. Australia.
Subgenus
Ropalichrus Allsopp, 1982.
Liparochrus
(Ropalichrus) monteithi Paulian, 1980. Australia.
*Liparochrus
(Ropalichrus) storeyi Paulian, 1980. Australia.
Pachyplectrinae
Ocampo
Type
species Pachyplectrus LeConte, 1874.
Pachyplectrus
LeConte, 1874 (1).
*Pachyplectrus
laevis LeConte, 1874. U.S.A.
Brenskea
Reitter, 1891 (2).
Spermohybosorus
Pic 1922. (synonym).
Brenskea chudeaui Reitter, 1909. Algeria.
*Brenskea
coronata Reitter, 1891. Central Asia, Iran, Israel, Morocco, Algeria,
Tunisia, Libya,
Egypt, Azerbaijan, Russia.
Brenskea varentzovi
Semenov, 1896 (synonym)
Spermohybosorus testaceous Pic, 1922 (synonym).
Incertae
sedis
Borrochrus
Allsopp, 1979 (2).
Trichops
de Borre, 1886. (preoccupied homonym). (type species Trichops ciliatus
de Borre, nec Val, 1853).
*Borrochrus
ciliatus (de Borre, 1886). (Trichops). Brazil, Paraguay,
Argentina.
Borrochrus
mutilus (Petrovitz, 1968). (Trichops). Paraguay.
Daimothoracodes Petrovitz,
1970 (4).
Daimothoracodes
confossus Ocampo and Vaz-de-Mello, 2002. Brazil.
Daimothoracodes
magnificus Martínez, 1994. Bolivia, Argentina.
*Daimothoracodes
mirabilis Petrovitz, 1970. Brazil.
Daimothoracodes
rugomarginatus Ocampo, 2005. Brazil.