HYBOSORIDAE Erichson, 1847

Anaidinae Nikolajev, 1996

Cryptogeniinae Howden, 2001 (new synonym).
Type genus Anaides Westwood, 1845.

Anaides Westwood, 1845 (13).

Aneides (Incorrect subsequent spelling by Balthasar 1838).

Anaides carioca Ocampo Brazil.

*Anaides fossulatus Westwood 1846. Panama, Colombia, Venezuela, Guyana, Trinidad and Tobago, Suriname.

Anaides reticulatus, Endrödi, 1962. (synonym).

Anaides howdeni Ocampo,  Dominican Republic.

Anaides laticollis Harold, 1863. Belize, Costa Rica, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, Panama.

Anaides longeciliatus Balthasar, 1938. Costa Rica, Panama.

Anaides onofrii Ocampo, Colombia, Ecuador, Peru, Bolivia.

Anaides ortii Ocampo, Venezuela.

Anaides parvulus Ocampo, Venezuela.

Anaides planus Ocampo, Costa Rica, Ecuador.

Anaides quinckei Ocampo Bolivia.

Anaides rugosus Robinson, 1948a. Ecuador, Peru.

Anaides simplicicollis Bates, 1887. Costa Rica, Panama.

Anaides vartorellii Ocampo Barbados.

 

Callosides Howden, 1971 (3).

*Callosides campbelli Howden, 1971. Colombia.

Callosides bartolozzii Paulian and Cambefort, 1995. Ecuador.

Callosides genieri Howden, 2001. Ecuador.

 

Chaetodus Westwood, 1845 (33).

Subgenus Chaetodus Westwood, 1845.

Pseudohybosorus Endrödi, 1963. (synonym).

Chaetodus allsoppi Martínez, 1988. Peru.

Chaetodus amazonicus de Borre, 1886. Brazil, Suriname, Venezuela.

            Pseudohybosorus drifti Endrödi, 1963 (synonym).

Chaetodus amazonicus inesperatus Martínez, 1988 (synonym).

Chaetodus bolivianus Martínez, 1956. Bolivia.

Chaetodus columbicus Petrovitz, 1970. Colombia.

Chaetodus datoi Ocampo, Bolivia

Chaetodus exaratus Arrow, 1909. Brazil, Argentina, Brazil, Paraguay.

Chaetodus apicipennis Petrovitz, 1970. (synonym).

Chaetodus fraternus Martínez, 1994. Bolivia.

Chaetodus globosus Ocampo, Mexico.

Chaetodus humerosus Petrovitz, 1970. Venezuela.

Chaetodus hoffmanni Ocampo, Panama, Venezuela.

Chaetodus irregularis Westwood, 1846. Bolivia, Brazil, Paraguay, Argentina, Uruguay.

Chaetodus striatus de Borre, 1886b (synonym).

Chaetodus jamesonae Ocampo, Venezuela.

Chaetodus lacandonicus Martínez and Morón, 1990. Mexico, Guatemala.

Chaetodus maquipucuna Ocampo, Ecuador.

Chaetodus mimi Ocampo, Ecuador, Peru, Bolivia, Brazil.

Chaetodus nigrifrons Ocampo, Ecuador.

Chaetodus noirregularis Ocampo, Brazil.

Chaetodus paucarae Ocampo, Ecuador.

Chaetodus paulseni Ocampo, Brazil.

Chaetodus pax Ocampo, Venezuela.

*Chaetodus piceus Westwood, 1846. Argentina, Bolivia, Brazil, Paraguay.

Chaetodus platynotus Ocampo, Ecuador.

Chaetodus ratcliffei Ocampo, Brazil.

Chaetodus rodolfo Ocampo, Ecuador.

Chaetodus sagittarius Ocampo, Brazil.

Chaetodus smithi Ocampo, Peru, Bolivia, Brazil.

Chaetodus teamscaraborum Ocampo Belize, Colombia, Costa Rica,

Guatemala, Honduras, Nicaragua, Panama, Venezuela.

Chaetodus villosicollis Benderitter, 1923. Argentina, Paraguay.

Subgenus Chaetodopsis Martínez, 1988.

*Chaetodus asuai Martínez, 1956. Colombia, Ecuador, Peru, Bolivia.

            Chaetodus brancuccii Martínez, 1994. Venezuela.

            Chaetodus octocarinatus Ocampo, Brazil, Guyana, Suriname.

            Chaetodus tricarinatus Ocampo, Colombia, Ecuador, Peru.

            Chaetodus venezolanus Martínez, 1994. Venezuela.

 

Cretanaides Nikolajev, 1996 (1).

*Cretanaides trogopterus Nikolajev, 1996. Siberia (Russia).

 

Cryptogenius Westwood, 1845 (2).

Cremastochilodius Krikken, 1975. (synonym, type species Cremastochilodius trisits Krikken 1975).

*Cryptogenius miersianus Westwood, 1846. Costa Rica, Panama, Colombia, Ecuador.

Cryptogenius fryi Arrow, 1909. Brazil, Argentina.

Cremastochilodius tristis Krikken, 1975. (synonym).

 

 Hybochaetodus Arrow, 1909 (2).

Hybochaetodus flaco Ocampo, 2002. Peru.

*Hybochaetodus obscurus Arrow, 1909. Peru.

 

Totoia Ocampo, 2003 (3).

Totoia brachycarina Ocampo, 2003. Nicaragua, Costa Rica. Panama.

*Totoia splendida Ocampo, 2003. Panama, Colombia.

Totoia magnifica Ocampo, Panama. 

 

Ceratocanthinae Martínez, 1968.

 Acanthocerinae Lacordaire, 1856 (synonym, see below under Ceratocanthus).

 Tribe Ceratocanthini Martínez, 1968.

 Acanthocerodes Péringuey, 1901 (3).

Acanthocerodes endroedyi Paulian, 1980. South Africa.

Acanthocerodes martini Paulian, 1992. South Africa.

*Acanthocerodes singularis Péringuey, 1901. South Africa.

 

Afrocloetus Petrovitz, 1968 (1).

Afrocloeotus (incorrect subsequent spelling by Paulian, 1977a)

*Afrocloetus gibbosus Petrovitz, 1968. Tanzania.

 

Aneilobolus Hesse, 1948 (4).

Aneilobolus endroedyyoungai Paulian, 1983. South Africa.

Aneilobolus gigas Paulian, 1977a. South Africa.

*Aneilobolus lawrencei Hesse, 1948. South Africa.

Aneilobolus leleupi Paulian, 1977. South Africa.

 

Anopsiostes Paulian, 1982 (1)

 *Anopsiostes punctatus Paulian, 1982. Ecuador, Peru.

 

Astaenomoechus Martínez and Pereira, 1959 (22).

Belloanopsiostes Vaz de Mello, 1996. (synonym). (type species: B. gilli Vaz de Mello, 1996).

Astaenomoechus americanus (Boucomont, 1936). (Philharmostes). Colombia, Costa Rica, Ecuador, French Guiana, Mexico, Peru, Trinidad.

Astaenomoechus termitophilus Petrovitz, 1976.  (synonym).

Astaenomoechus andersoni Howden and Gill, 2000. Venezuela.

Astaenomoechus cavei Howden and Gill, 2003. Belize, Honduras.

Astaenomoechus cicheroi Martínez, 1969. Bolivia.

Astaenomoechus criberrimus Paulian, 1982. Peru.

Astaenomoechus estriatus Paulian, 1982. Ecuador.

Astaenomoechus gilli (Vaz de Mello, 1996). (Belloanopsiostes). Brazil.

Astaenomoechus hispidus Howden and Gill, 2003. Costa Rica.

*Astaenomoechus hospes (Wasmann, 1902). (Sphaeromorphus). Argentina, Brazil.

Astaenomoechus luniferus Petrovitz, 1973. Colombia, Ecuador.

Astaenomoechus mixtus Howden and Gill, 2003. Costa Rica, Panama.

Astaenomoechus multipunctatus Howden and Gill, 2003. Nicaragua, Costa Rica, Panama.

Astaenomoechus nevermanni (Boucomont, 1936). (Philharmostes). Costa Rica.

Astaenomoechus redtenbacheri (Harold, 1874). (Acanthocerus). Colombia, Venezuela.

Astaenomoechus paniculus Howden and Gill, 2003. Costa Rica, Panama.

Astaenomoechus parvosetosus Howden and Gill, 2003. Costa Rica, Panama.

Astaenomoechus punctifrons Howden and Gill, 2003. Nicaragua, Costa Rica.

Astaenomoechus setosus (Boucomont, 1936). (Philharmostes). Brazil, Colombia, Costa Rica, Ecuador, Guyana.

Astaenomoechus setulosus (Harold, 1874). (Acanthocerus). Colombia, French Guiana, Venezuela.

Astaenomoechus solisi Howden and Gill, 2003. Nicaragua, Costa Rica.

Astaenomoechus strigulosus Howden and Gill, 2003. Costa Rica

Astaenomoechus unidentatus Petrovitz, 1973. Colombia, Ecuador.

 

Aulisostes Howden and Gill, 2000 (2).

Aulisostes paradoxus (Paulian, 1982). (Germarostes). Brazil.

*Aulisostes pseudoparadoxus Howden and Gill, 2001. Colombia.

 

Baloghianestes Paulian, 1968 (1).

*Baloghianestes lissoubai Paulian, 1968. Cameroon, Congo.

 

Besuchetostes Paulian, 1972 (11).

Besuchetostes besucheti (Paulian, 1972). (Cyphopisthes). Sri Lanka.

Philharmostes carinatus Paulian, 1975. (synonym).

Besuchetostes dubium Paulian, 1975. India.

Besuchetostes hindu Paulian, 1975. India.

Besuchetostes howdeni Paulian, 1979. Malaysia.

Besuchetostes jaccoudi Paulian, 1977. Malaysia.

Besuchetostes keralae Paulian, 1975. India.

Besuchetostes kodaikanalense Paulian, 1975. India.

Besuchetostes loebli Paulian, 1972. Sri Lanka.

Besuchetostes mussardi Paulian, 1972. Sri Lanka.

Besuchetostes peradeniyae Paulian, 1972. Sri Lanka.

*Besuchetostes taprobanae Paulian, 1972. Sri Lanka.

 

Callophilharmostes Paulian, 1968 (1).

Philharmostes (Callophilharmostes) Paulian, 1968

*Callophilharmostes fleutiauxii (Paulian, 1943). (Philharmostes). Cameroon, Gabon, Guinea,

Ivory Coast.

 

Carinophilharmostes Paulian, 1968 (1).

Philharmostes (Carinophilharmostes) Paulian, 1968.

*Carinophilharmostes vadoni (Paulian, 1937). (Philharmostes). Cameroon, Central African

 Republic, Congo, Zaire, Equatorial Guinea, Guinea, Ivory Coast, Uganda.

Pterorthochaetes multigibber Petrovitz, 1967. (synonym).

 

Ceratocanthoides Paulian, 1982 (1).

*Ceratocanthoides undatus (Petrovitz, 1973). (Acanthocerus). Brazil, Colombia, Peru.

 

Ceratocanthopsis Paulian, 1982 (3).

*Ceratocanthopsis fulgida (Martínez, 1967). (Acanthocerus). Brazil.

Ceratocanthopsis pernitida Paulian, 1982. Bolivia, Brazil, French Guiana, Trinidad.

Ceratocanthopsis pygmaea (Harold, 1874). (Acanthocerus). Brazil, French Guiana, Venezuela, Suriname.

 

Ceratocanthus White, 1842 (53).

Acanthocerus MacLeay, 1819. (synonym). (junior homonym of Acanthocerus Palisot de Beauvois, 1818, Hemiptera)

Gymnoropterus Gestro, 1899. (synonym). (type species Synarmostes striatulus Lansberge, 1887).

Sphaeromorphus Germar, 1843. (synonym).

*Ceratocanthus aeneus (MacLeay, 1819). (Acanthocerus). USA.

Sphaeromorphus volvox Erichson in Germar, 1843 (synonym).

Ceratocanthus amazonicus Paulian, 1982. Brazil, Colombia, French Guiana.

Ceratocanthus aureolus (Harold, 1874). (Acanthocerus). Brazil.

Ceratocanthus baniensis Howden, 1978. Hispaniola.

Ceratocanthus basilicus (Germar, 1843). (Sphaeromorphus). Brazil.

Ceratocanthus bicinctoides Paulian, 1982. Brazil.

Ceratocanthus bicinctus (Germar, 1843). (Sphaeromorphus). Brazil.

Ceratocanthus bonfilsi Chalumeau, 1977. Guadeloupe.

Ceratocanthus brasiliensis (Lansberge, 1887). (Acanthocerus). Brazil.

Ceratocanthus chalceus (Germar, 1843). (Sphaeromorphus). Cuba.

Ceratocanthus clypealis (Lansberge, 1887). (Acanthocerus). Brazil, French Guiana, Peru.

Ceratocanthus ebeninus (Erichson in Germar, 1843). (Sphaeromorphus). Brazil.

Ceratocanthus eberti Paulian, 1982. Brazil.

Ceratocanthus eulampros (Bates, 1887). (Acanthocerus). Costa Rica, Nicaragua, Panama.

Ceratocanthus fuscoviridis (Ohaus, 1911). (Acanthocerus). Argentina.

Ceratocanthus globulus (Erichson in Germar, 1843). (Sphaeromorphus). Brazil, Colombia, French Guiana.

Ceratocanthus gundlachi (Harold, 1874). (Acanthocerus). Cuba.

Ceratocanthus humeralis (Erichson in Germar, 1843)). (Sphaeromorphus). Brazil, Panama, Suriname.

Ceratocanthus inca Paulian, 1982. Peru.

Ceratocanthus major Paulian, 1982. French Guiana.

Ceratocanthus mathani Paulian, 1982. Peru.

Ceratocanthus micans (Harold, 1874). (Acanthocerus). Brazil.

Ceratocanthus micros (Bates, 1887). (Acanthocerus). Costa Rica, Mexico.

Ceratocanthus monrosi (Martínez and Pereira, 1959). Bolivia, Brazil.

Ceratocanthus nanus (Germar, 1843). (Sphaeromorphus). Brazil, Paraguay.

Ceratocanthus niger Paulian, 1982b. Brazil.

Ceratocanthus nitidus (Germar, 1843). (Sphaeromorphus). Bolivia, Brazil.

Ceratocanthus pararelucens Howden, 1978. St. Vincent.

Ceratocanthus pauliani (Delgado and Hernandez, 1998). (Anopsiostes). Mexico.

Ceratocanthus pecki Paulian, 1982. Ecuador.

Ceratocanthus perpunctatus Paulian, 1982. Ecuador, Peru.

Ceratocanthus politus (Erichson in Germar, 1843). (Sphaeromorphus). Argentina, Bolivia, Brazil, Peru.

Ceratocanthus pseudosuturalis Paulian, 1982. Brazil.

Ceratocanthus punctolineatus Paulian, 1982. Brazil, Colombia.

Ceratocanthus punctulatus (Lansberge, 1887). (Acanthocerus). Brazil, Peru.

Ceratocanthus pyritosus (Erichson in Germar, 1843). (Sphaeromorphus). St. Thomas.

Ceratocanthus quadristriatus Paulian and Vaz de Mello, 1998. Brazil.

Ceratocanthus relucens mexicanus (Bates, 1887). (Acanthocerus). Belize, Mexico.

Ceratocanthus relucens relucens (Bates, 1887). (Acanthocerus). Belize, Costa Rica, Guatemala, Mexico, Nicaragua, Panama.

Ceratocanthus rotundicollis (Bates, 1887). (Acanthocerus). Panama.

Ceratocanthus semipunctatus carioca (Martínez, 1967). (Acanthocerus). Brazil.

Ceratocanthus semipunctatus semipunctatus (Germar, 1843). (Sphaeromorphus). Brazil, Colombia, Paraguay.

Ceratocanthus semistriatus (Germar, 1843). (Sphaeromorphus). Cuba.

Ceratocanthus seriatus (Erichson in Germar, 1843). (Sphaeromorphus). Brazil, French Guiana, Peru.

Ceratocanthus sesquistriatus (Germar, 1843). (Sphaeromorphus). Brazil.

Ceratocanthus sexstriatus Paulian, 1982. Brazil.

Ceratocanthus spinicornis (Fabricius, 1792). (Trox). Unknown distribution.

Ceratocanthus steinbachi Paulian, 1982b. Bolivia.

Ceratocanthus striatulus (Lansberge, 1887). (Synarmostes). Unknown distribution.

Ceratocanthus suturalis (Lansberge, 1887). (Acanthocerus). Bolivia, Brazil, Ecuador, French Guiana, Peru, Surinam, Trinidad.

Ceratocanthus suturaloides Paulian, 1982. Venezuela.

Ceratocanthus termiticola (Wasmann, 1894). (Acanthocerus). Bolivia, Brazil, Colombia, Ecuador.

Ceratocanthus turquinensis (Zayas, 1988). (Acanthocerus). Cuba.

Ceratocanthus undulatus (Harold, 1874). (Acanthocerus). Brazil, French Guiana.

Ceratocanthus vicarius (Bates, 1887). (Acanthocerus). Belize, Guatemala, Mexico.

 

Chaetophilharmostes Paulian, 1977 (1).

*Chaetophilharmostes chevalieri (Paulian, 1937). (Philharmostes). Congo, Guinea, Ivory Coast,

Liberia.

Pterorthochaetes termitophilus Petrovitz, 1968. (synonym).

 

Cloeotus Germar, 1843 (3).

*Cloeotus latebrosus Germar, 1843. Colombia.

Cloeotus petrovitzi Paulian, 1982. Brazil.

Cloeotus semicostatus Germar, 1843. Colombia.

 

Congomostes Paulian, 1968 (2).

*Congomostes baloghi Paulian, 1968. Congo (=former Zaire.).

Congomostes janssensi (Basilewsky, 1955). (Philharmostes). Congo (=former Zaire).

 

Cryptophilharmostes Ballerio, 2000 (2).

*Cryptophilharmostes mahunkai Ballerio, 2000. Tanzania.

            Cryptophilharmostes merkli Ballerio, 2005. Tanzania

 

Cyphopisthes Gestro, 1899 (10).

*Cyphopisthes amphicyllis (Sharp, 1875). (Synarmostes). Indonesia, "New Guinea."

Cyphopisthes crux  (Sharp, 1875). (Synarmostes). "Borneo."           

Cyphopisthes descarpentriesi Paulian, 1977c. Australia.

Cyphopisthes dohertyi Paulian, 1942. (Philharmostes). India.

Cyphopisthes inexpectatus Paulian, 1981. Philippines.

Cyphopisthes krikkeni Paulian, 1980. Indonesia.

Cyphopisthes luzonicus Paulian, 1978. Philippines.

Cyphopisthes minutus Paulian, 1978. Malaysia.

Cyphopisthes szentivanyi (Paulian, 1973). (Philharmostes). Papua New Guinea.

Cyphopisthes wallacei (Pascoe, 1860). (Sphaeromorphus). Malaysia, Indonesia.

                        Cyphopisthes humeralis Gestro, 1899 (synonym).

 

Ebbrittoniella Martínez, 1962 (2).

Eubrittoniella (incorrect subsequent spelling by Paulian, 1978)

Ebbrittoniella gestroi (Paulian, 1942). (Philharmostes). Indonesia, Malaysia,

*Ebbrittoniella ignita (Westwood, 1883). (Acanthocerus (Sphaeromorphus). Indonesia, Malaysia.

 

Eusphaeropeltis Gestro, 1899 (16).

Eusphaeropeltis aureola Gestro, 1899. Indonesia (Sumatra).

*Eusphaeropeltis aurora (Lansberge, 1887). (Synarmostes). Indonesia (Sumatra).

Eusphaeropeltis borneensis Paulian, 1978. "Borneo."

Eusphaeropeltis boucomonti Paulian, 1978. Indonesia (Sumatra).

Eusphaeropeltis celebicus Paulian, 1978. Indonesia (Sulawesi).

Eusphaeropeltis corruscus Gestro, 1899. Malaysia.

Eusphaeropeltis drescheri Paulian, 1942. Indonesia (Java).

Eusphaeropeltis iris Gestro, 1899. Indonesia (Sumatra).

Eusphaeropeltis kedahensis Paulian, 1942. Indonesia (Sumatra).

Eusphaeropeltis krikkeni Paulian, 1978. Indonesia (Sumatra).

Eusphaeropeltis modiglianii Gestro, 1899. Indonesia (Sumatra).

Eusphaeropeltis pulcher (Lansberge, 1885). (Synarmostes). Indonesia (Sumatra).

Eusphaeropeltis punctatissimus (Lansberge, 1887). (Synarmostes). Brunei, Malaysia.

Eusphaeropeltis raapi Gestro, 1899. Indonesia (Nias).

Eusphaeropeltis sabah Paulian, 1989. Malaysia (Sabah).

Eusphaeropeltis scheuerni Paulian, 1982a. Indonesia (Sumatra).

 

Germarostes Paulian, 1982 (69).

Subgenus Germarostes Paulian, 1982.

Germarostes (Germarostes) allorgei (Paulian, 1947). (Cloeotus). Dominica, Guadeloupe.

Germarostes (Germarostes) anchicayae Paulian, 1982. Colombia.

Germarostes (Germarostes) antiquus (Erichson in Germar, 1843). (Acanthocerus). Peru, Surinam, Venezuela.

*Germarostes (Germarostes) aphodioides aphodioides  (Illiger, 1800). (Melolontha). Canada to Argentina.

Acanthocerus laevistriatus  Laporte, 1840. (synonym).

            Scarabaeus latipes Germar, 1824. (synonym).

Trox splendidus Say, 1835. (synonym).

Germarostes (Germarostes) aphodioides prionomus (Bates, 1887). (Cloeotus). Guatemala.

Germarostes (Germarostes) argentinus (Ohaus, 1911). (Cloeotus). Argentina.

Germarostes (Germarostes) bidens (Bates, 1887). (Cloeotus). Brazil, Panama.

Germarostes (Germarostes) carinatus Paulian, 1982. Ecuador.

Germarostes (Germarostes) carltoni Howden & Gill, 2005. Peru.

Germarostes (Germarostes) costulatus Paulian, 1982. Colombia.

Germarostes (Germarostes) crassicollis (Arrow, 1903). (Cloeotus). St. Vincent.

Germarostes (Germarostes) degallieri Paulian, 1982. Colombia, Ecuador, French Guiana.

Germarostes (Germarostes) ecuadoricus Paulian, 1982. Ecuador.

Germarostes (Germarostes) excisus (Bates, 1887). (Cloeotus). Guatemala, Mexico.

Germarostes (Germarostes) farri (Howden, 1970). (Cloeotus).  Jamaica.

Germarostes (Germarostes) gaujoni Paulian, 1982. Ecuador.

Germarostes (Germarostes) globosus (Say, 1835). (Trox). Mexico, USA.

Germarostes (Germarostes) heterodynamus Paulian, 1982. Brazil.

Germarostes (Germarostes) howdenicus Paulian, 1982. Ecuador.

Germarostes (Germarostes) infantulus (Bates, 1887). (Cloeotus). Guatemala.

Germarostes (Germarostes) instriatus Paulian, 1982. Ecuador.

Germarostes (Germarostes) jamaicensis (Howden, 1978). (Cloeotus). Jamaica.

Germarostes (Germarostes) leticiae Paulian, 1982.. Brazil, Colombia.

Germarostes (Germarostes) macleayi (Perty, 1830). (Acanthocerus). Guatemala, Panama, Colombia, French Guiana, Peru, Brazil, Bolivia, Argentina, Paraguay.

Germarostes (Germarostes) oberthueri Paulian, 1982. Brazil.

Germarostes (Germarostes) pauliani (Chalumeau  and Cambefort, 1976). (Cloeotus). Guadaloupe

Germarostes (Germarostes) pecki (Howden, 1970). (Cloeotus). Jamaica.

Germarostes (Germarostes) plicatus (Erichson in Germar, 1843)). (Acanthocerus). Mexico to

Argentina except Chile.

Germarostes (Germarostes) posticus (Germar, 1843). (Acanthocerus). Argentina?, Chile,

Acanthocerus muricatus Curtis, 1845. (synonym).

Cloeotus posticus mochae Gutiérrez, 1946 (synonym?).

Germarostes (Germarostes) pullus Paulian, 1982b. Ecuador

Germarostes (Germarostes) puncticollis (Erichson in Germar, 1843). (Acanthocerus). Brazil,

Ecuador, Paraguay.

Germarostes (Germarostes) punctulatus (Ohaus, 1911). (Cloeotus). Argentina, Bolivia, Brazil,

Colombia, French Guiana, Paraguay.

Germarostes (Germarostes) pusillus ( Laporte, 1840). (Acanthocerus). Colombia.

Germarostes (Germarostes) pustulosus (Lansberge, 1887). (Cloeotus). Colombia.

Germarostes (Germarostes) reticularis (Bates, 1887). (Cloeotus). Mexico.

Germarostes (Germarostes) rotundatus Paulian, 1982. Venezuela.

Germarostes (Germarostes) rufopiceus (Arrow, 1903). (Cloeotus). St. Lucia, St. Vincent.

Germarostes (Germarostes) rugatus (Germar, 1843). (Acanthocerus). Colombia.

Acanthocerus rugosus Germar 1843 (synonym).

Germarostes (Germarostes) salesiacus Paulian, 1982. Brazil.

Germarostes (Germarostes) semituberculatus (Germar 1843). (Acanthocerus). Bolivia, Brazil,

Paraguay, Peru, Venezuela.

Cloeotus acuarius Petrovitz, 1973. (synonym).

Germarostes (Germarostes) sinuatus sejunctus (Bates, 1887). (Cloeotus). Guatemala.

Germarostes (Germarostes) sinuatus sinuatus (Bates, 1887). (Cloeotus). Belize, Guatemala, Mexico.

Germarostes (Germarostes) sticticus (Erichson in Germar, 1843). (Acanthocerus). Brazil, French Guiana, Nicaragua, Suriname.

Germarostes (Germarostes) strigilateris (Bates, 1887). (Cloeotus). Mexico.

Germarostes (Germarostes) sulcipennis (Harold, 1875). (Cloeotus). Peru.

Germarostes (Germarostes) viridis (Lansberge, 1887). (Cloeotus). Mexico.

Subgenus Haroldostes Paulian, 1982b.

Germarostes (Haroldostes) abruptus (Petrovitz, 1973). (Cloeotus). Brazil.

Germarostes (Haroldostes) batesi (Harold, 1874a). (Cloeotus). Brazil, Bolivia.

Germarostes (Haroldostes) brunnipes (Germar, 1843). (Acanthocerus). Bolivia, Ecuador, Paraguay.

Germarostes (Haroldostes) bugabensis (Arrow, 1903). (Cloeotus). Panama.

Germarostes (Haroldostes) columbianus Paulian, 1982. Colombia.

Germarostes (Haroldostes) diffundus (Petrovitz, 1976). (Cloeotus). Brazil, Ecuador, Paraguay.

Germarostes (Haroldostes) ecuadoriensis (Petrovitz, 1976). (Cloeotus). Ecuador, Venezuela.

Germarostes (Haroldostes) geayi Paulian, 1982. Peru.

Germarostes (Haroldostes) guyanensis Paulian, 1982b. French Guiana, Panama?

Germarostes (Haroldostes) hamiger (Ohaus, 1911). (Cloeotus). Argentina, Bolivia.

Germarostes (Haroldostes) haroldi (Arrow, 1911). (Cloeotus). Colombia, Venezuela.

Cloeotus puncticollis Harold, 1874. (synonym) (homonym of Germarostes (Germarostes) puncticollis Erichson in Germar, 1843).

Germarostes (Haroldostes) indigaceus (Germar, 1843). (Acanthocerus). Brazil, Colombia, Ecuador, Venezuela.

Germarostes (Haroldostes) leprieuri (Germar, 1843). (Acanthocerus). Brazil, Ecuador, French Guiana.

Germarostes (Haroldostes) madeiranus Paulian, 1982b. Brazil.

Germarostes (Haroldostes) malkini Paulian, 1982b. Brazil.

Germarostes (Haroldostes) metallicus (Harold, 1874a). (Cloeotus). Brazil, Colombia, Ecuador, French Guiana.

Germarostes (Haroldostes) nasutus (Bates, 1887). (Cloeotus). Mexico.

Germarostes (Haroldostes) nigerrimus (Blanchard, 1841). (Sphaerelytrus). Bolivia, Colombia, Ecuador.

 Cloeotus grandis Petrovitz, 1973. (synonym).

Germarostes (Haroldostes) nitens (Guérin-Méneville, 1839). (Acanthocerus). Costa Rica, Brazil,

Honduras, Mexico, Panama.

Cloeotus acutipes Arrow, 1903. (synonym).

*Germarostes (Haroldostes) rugiceps (Germar, 1843). (Acanthocerus). Brazil, Panama?, Paraguay.

Germarostes (Haroldostes) senegalensis (Laporte, 1840). (Acanthocerus). Brazil, French Guiana.

Acanthocerus striatus Germar, 1843. (synonym).

Germarostes (Haroldostes) tibialis (Petrovitz, 1973). (Cloeotus). Brazil.

Germarostes (Haroldostes) tubericauda (Bates, 1891). (Cloeotus). Colombia, Ecuador.

Germarostes (Haroldostes) viridipennis (Bates, 1887). (Cloeotus). Costa Rica, Panama.

Germarostes (Haroldostes) viridulus (Bates, 1887). (Cloeotus). Mexico.

 

Glyptogermarostes (1)

Glyptopterus Paulian, 1982 (synonym, homonym of Glyptopterus de Chaudoir, 1838 Carabidae).

*Glyptogermarostes oberthueri (Paulian, 1982). (Glyptopterus). Brazil.

 

Goudotostes Paulian, 1979 (1).

*Goudotostes scabrosus ( Laporte, 1840). (Acanthocerus). Madagascar.

 

Macrophilharmostes Paulian, 1978 (1).

*Macrophilharmostes major (Paulian, 1975a). (Cyphopishtes). Papua New Guinea

 

Madrasostes Paulian, 1975 (28).

Madrasostes agostii Paulian, 1993. Malaysia.

Madrasostes boucomonti Paulian, 1978. Indonesia, Malaysia.

Madrasostes burckhardti Paulian, 1989. Malaysia.

Madrasostes clypeale Paulian, 1993. Indonesia (Sumatra).

Madrasostes depressum Paulian, 1992. Indonesia (Sumatra).

Madrasostes feae Gestro, 1899. India, Kampuchea, Myanmar, Nepal, Thailand.

Madrasostes franzi Paulian, 1978. Thailand.

Madrasostes granulatum (Paulian, 1975). (Cloeotus). Papua New Guinea.

Madrasostes inaequale Paulian, 1992. Indonesia (Sumatra).

Madrasostes kazumai hisamatsui Ochi, 1990. Japan.

Madrasostes kazumai kazumai Ochi, Johki, and Nakata, 1990. Japan.

Madrasostes loebli Paulian, 1981b. Papua New Guinea (Bismarck Isl.).

Madrasostes malayanum Paulian, 1979. Malaysia.

Madrasostes masumotoi Paulian, 1987. Thailand.

*Madrasostes nigrum Paulian, 1975. India.

Madrasostes orousseti Paulian, 1981. Philippines.

Madrasostes parcepunctatum Paulian, 1989. Malaysia.

Madrasostes punctatum Paulian, 1989. Malaysia.

Madrasostes rafflesi Paulian, 1979c. Malaysia.

Madrasostes reticulatum (Lansberge, 1887). (Synarmostes). Indonesia (Sulawesi), Philippines.

Madrasostes sabah Paulian, 1989. Malaysia (Sabah).

Madrasostes sculpturatum Paulian, 1989. Indonesia, Malaysia.

Madrasostes simplex Paulian, 1989. Malaysia.

Madrasostes sumatranum Paulian, 1992a. Indonesia (Sumatra).

Madrasostes tamil (Paulian, 1975b). (Philharmostes). India.

Madrasostes thai Paulian, 1987. Thailand.

Madrasostes thoracicum Paulian, 1989. Malaysia.

Madrasostes tonkinense (Paulian, 1945). (Cloeotus). Vietnam.

Madrasostes variolosum (Harold, 1874b). (Cloeotus). Indonesia, Malaysia.

 

Martinezostes Paulian, 1982 (3).

*Martinezostes asper (F. Philippi, 1859). (Acanthocerus). Chile.

Martinezostes fortecostatus (Gutiérrez, 1949). (Cloeotus). Chile.

Martinezostes ruizi (Gutiérrez, 1946). (Cloeotus). Chile.

 

Melanophilharmostes Paulian, 1968  (17).

Philharmostes (Melanophilharmostes) Paulian, 1968

Melanophilharmostes ashantii (Paulian, 1974). (Philharmostes). Ghana.

Melanophilharmostes bicarinatus (Paulian, 1974). (Philharmostes). Ghana.

Melanophilharmostes burgeoni (Paulian, 1946). (Pterorthochaetes). Cameroon, Zaire, Angola?

Melanophilharmostes carinatus (Paulian, 1974). (Philharmostes). Ghana.

Melanophilharmostes carvalhoi (Martínez, 1970). (Astaenomoechus). Angola.

Melanophilharmostes demirei Paulian, 1977. Cameroon.

Melanophilharmostes donisi (Basilewsky, 1955). (Pterorthochaetes). Zaire.

Melanophilharmostes endroedyi (Paulian, 1968. (Philharmostes (Melanophilharmostes) Zaire.

Melanophilharmostes ghanae (Paulian, 1974). (Philharmostes). Ghana, Togo.

Melanophilharmostes ocellatus (Paulian, 1968). (Philharmostes (Melanophilharmostes). Congo.

Melanophilharmostes palustris (Petrovitz, 1968). (Philharmostes). Zaire.

Melanophilharmostes puncticeps (Paulian, 1946). (Pterorthochaetes). Zaire, Liberia.

Melanophilharmostes posthi (Paulian, 1937). (Pterorthochaetes). Ivory Coast, Liberia, Togo.

Melanophilharmostes pseudoposthi (Paulian, 1977). Zaire.

Melanophilharmostes pygmaeus (Petrovitz, 1968). (Philharmostes). Zaire.

Melanophilharmostes vincenti (Paulian, 1968). (Philharmostes (Melanophilharmostes). Congo.

*Melanophilharmostes zicsii (Paulian, 1968). (Philharmostes (Melanophilharmostes). Cameroon, Congo, Gabon.

 

Nesopalla Paulian and Howden, 1982 (2).

Nesopalla borinquensis Paulian and Howden, 1982. Puerto Rico.

*Nesopalla iviei Paulian and Howden, 1982. Puerto Rico, Virgin Islands.

 

Paulianostes Ballerio, 2000 (3).

Paulianostes acromialis (Pascoe, 1860). (Sphaeromorphus). Malaysia.

Philharmostes arrowi Paulian, 1942 (synonym).

*Paulianostes georyssoides (Gestro, 1899). (Cyphopisthes). Indonesia, Malaysia.

Paulianostes panggoling Ballerio, 2000. Brunei, Malaysia (Sabah).

 

Perignamptus Harold, 1877 (4 ).

Perignamptus carinipennis Gestro, 1899. Papua New Guinea.

Perignamptus loriae Gestro, 1899. Papua New Guinea.

Perignamptus rossi Paulian, 1978. Indonesia (Irian Jaya).

*Perignamptus sharpi Harold, 1877. Papua New Guinea.

 

Petrovitzostes Paulian, 1977  (1).

*Petrovitzostes guineensis (Petrovitz, 1968). (Pterorthochaetes). Cameroon, Equatorial Guinea.

 

Philharmostes Kolbe, 1895 (31).

Subgenus Philharmostes Kolbe, 1895.

Philharmostes (Philharmostes) adami Paulian, 1968. (Philharmostes (Carinophilharmostes). Congo, Guinea, Ivory Coast.

*Philharmostes (Philharmostes) aeneoviridis Kolbe, 1895. Madagascar.

Philharmostes (Philharmostes) basicollis (Fairmaire, 1897). Madagascar.

Philharmostes obscurus Fairmaire, 1900 (synonym).

Philharmostes (Philharmostes) basilevskyi Paulian, 1977. Tanzania.

Philharmostes (Philharmostes) bicolor Boucomont, 1937. Madagascar.

Philharmostes (Philharmostes) boucomonti Petrovitz, 1968. Madagascar.

Philharmostes (Philharmostes) caffer (Martínez, 1970) (Astaenomoechus). Angola.

Philharmostes (Philharmostes) corruscus Fairmaire, 1903. Madagascar.

Philharmostes pilula Fairmaire, 1903 (synonym).

Philharmostes convexifrons Fairmaire, 1903 (synonym).

Philharmostes (Philharmostes) criberrimus Paulian, 1979. Madagascar.

Philharmostes (Philharmostes) cribrarius Fairmaire, 1903. Madagascar.

Philharmostes (Philharmostes) cupreolus Fairmaire, 1900. Madagascar.

Philharmostes (Philharmostes) descarpentriesi Paulian, 1979. Madagascar.

Philharmostes (Philharmostes) disparilis Hesse, 1948. South Africa.

Philharmostes (Philharmostes) garciabesi (Martínez, 1970) (Astaenomoechus). Congo (= former Zaire)

Philharmostes (Philharmostes) girardi Paulian, 1993. Guinea.

Philharmostes (Philharmostes) grebennikovi Ballerio, 2004. Tanzania

Philharmostes (Philharmostes) integer Kolbe, 1895. Tanzania.

Philharmostes (Philharmostes) interruptus Hesse, 1948. South Africa.

Philharmostes (Philharmostes) latericostatus (Fairmaire, 1885). (Synarmostes). Madagascar.

Synarmostes obscuroaeneus Fairmaire, 1897 (synonym).

Synarmostes perroti Wasmann, 1897 (synonym).

Philharmostes (Philharmostes) olsoufieffi Boucomont, 1937. Madagascar.

Philharmostes (Philharmostes) ornatus Ballerio, 2004. Tanzania

Philharmostes (Philharmostes) perrieri (Fairmaire, 1898). (Synarmostes). Madagascar.

Philharmostes (Philharmostes) pseudobasicollis Paulian, 1979. Madagascar.

Philharmostes (Philharmostes) pseudumbratilis Ballerio, 2004. Tanzania

Philharmostes (Philharmostes) spelaeus Paulian, 1979. Zaire.

Philharmostes (Philharmostes) umbilicatus Petrovitz, 1968. Zaire, Guinea

Philharmostes (Philharmostes) umbratilis Petrovitz, 1968. Tanzania.

Philharmostes (Philharmostes) vadonianus Paulian, 1979. Madagascar.

Philharmostes (Philharmostes) werneri Ballerio, 2001. Tanzania.

Philharmostes (Philharmostes) zuluensis Hesse, 1948. South Africa.

Subgenus Holophilarmostes Paulian, 1968.

*Philharmostes (Holophilharmostes) badius (Petrovitz, 1967). (Pterorthochaetes). Cameroon,

Congo, Zaire, Guinea.

Philharmostes (Holophilharmostes) cohici Paulian, 1968. (synonym).

 

Pseudopterorthochaetes Paulian, 1977 (7).

Pseudopterorthochaetes cambeforti Paulian, 1981. Ivory Coast.

Pseudopterorthochaetes criberrimus Paulian, 1977. Zaire, Gabon.,

*Pseudopterorthochaetes elytratus (Paulian, 1946). (Pterorthochaetes). Cameroon, Zaire.

Pseudopterorthochaetes endroedyi (Paulian, 1974). (Pterorthochaetes). Ghana, Ivory Coast.

Pseudopterorthochaetes hystrix Paulian, 1991. Madagascar.

Pseudopterorthochaetes kumasii (Paulian, 1974). (Pterorthochaetes). Gabon, Ghana.

Pseudopterorthochaetes machadoi (Martínez, 1970). (Astaenomoechus). Angola.

 

Pterorthochaetes Gestro, 1899 (21).

Pterorthochaetes andamanus Paulian, 1937. India (Andaman Islands).

Pterorthochaetes armatus Paulian, 1945. Vietnam.

Pterorthochaetes brevis (Sharp, 1875). (Synarmostes). Indonesia (Irian Jaya), Papua New Guinea.

Acanthocerus (Sphaeromorphus) byrrhoides Westwood, 1883 (synonym).

Pterorthochaetes brevisetosus Gestro, 1899. Singapore.

Pterorthochaetes coomani Paulian, 1945. Vietnam.

Pterorthochaetes cribricollis Gestro, 1899. Australia. Papua New Guinea.

*Pterorthochaetes gestroi gestroi (Harold, 1874). (Synarmostes). Indonesia, Malaysia.

Pterorthochaetes gestroi longisetosus Gestro, 1899. Singapore.

Pterorthochaetes haroldi (Sharp, 1875). (Synarmostes). Indonesia, Malaysia, Singapore.

Pterorthochaetes hirtus Gestro, 1899. Indonesia (Sumatra).

Pterorthochaetes incertus Gestro, 1899. Malaysia.

Pterorthochaetes insularis Gestro, 1899. Indonesia, Malaysia, Nepal, Thailand.

Pterorthochaetes latus (Sharp, 1875). (Synarmostes). Indonesia (Sumatra), Singapore.

Pterorthochaetes lavongai Paulian, 1968b. Papua New Guinea (Bismarck Isl.).

Pterorthochaetes laxepunctatus Paulian, 1973. Papua New Guinea.

Pterorthochaetes mallicoloi Paulian, 1968b. Vanuatu Island.

Pterorthochaetes montanus Ballerio, 1999. Malaysia.

Pterorthochaetes picinus (Sharp, 1875). (Synarmostes). Micronesia, Philippines.

Pterorthochaetes puncticollis (Sharp, 1875). (Synarmostes). Indonesia (Java).

Pterorthochaetes septemtrionalis Ballerio, 1999. Nepal.

Pterorthochaetes simplex Gestro, 1899. Australia, Papua New Guinea.

Pterorthochaetes sulawesii Paulian 1987. Indonesia (Sulawesi).

 

 

Synarmostes Germar, 1843 (4).

            Synarmostes antsingyi Paulian, 1979. Madagascar.

Synarmostes humilis Fairmaire, 1893. Comores islands.

Synarmostes niger Paulian, 1979. Madagascar.

*Synarmostes tibialis (Klug, 1832). (Acanthocerus). Madagascar.

 

 

Tribe Ivieolini Howden and Gill, 2000.

 

Ivieolus Howden and Gill, 1988 (3).

Ivieolus brooksi Howden and Gill, 2000. French Guiana.

Ivieolus inflaticollis Howden and Gill, 2001. Ecuador.

*Ivieolus pseudoscutellatus Howden and Gill, 1988. Guyana, Venezuela.

 

 

Tribe Scarabatermitini Nikolajev, 1999

 

Scarabaeinus Silvestri, 1940  (1).

*Scarabaeinus termitophilus Silvestri, 1940. Brazil.

 

Scarabatermes Howden, 1973 (1).

*Scarabatermes amazonensis Howden, 1973. Colombia.

Trachycrusus Howden and Gill, 1995 (2).

*Trachycrusus lescheni Howden and Gill, 1995. Peru.
Trachycrusus striatulus Howden and Gill, 1995. Peru.

 

Xenocanthus Howden and Gill, 1988 (1).

*Xenocanthus singularis Howden and Gill, 1988. Venezuela.

 

Hybosorinae Erichson, 1847.

Type genus Hybosorus MacLeay, 1819.

 Apalonychus Westwood, 1845 (4).

Hapalonychus (incorrect subsequent spelling by Lacordaire 1856).

Haplonychus (incorrect subsequent spelling by Petrovitz (1973).

Trichops Val, 1853. (synonym). (type sepecies Trichops testaceus Val, 1853)

Apalonychus nattereri Petrovitz, 1973. Brazil.

Apalonychus pusillus Arrow, 1911. Paraguay.

Apalonychus rufulus ( Laporte, 1840). (Hybosorus). Dominican Republic.

*Apalonychus waterhousei Westwood, 1846. Cuba, Dominican Republic.

Trichops testaceus Val, 1853. (synonym)

Apalonychus rufulus de Borre, 1886. (synonym).

 

Araeotanypus Waterhouse, 1875 (6).

Araeotanopus (Incorrect subsequent spelling by Arrow (1912)).

Araeotanypus bicolor Petrovitz, 1967. South Africa.

*Araeotanypus boops Waterhouse, 1875. Botswana, South Africa.

Araeotanypus consors Péringuey, 1908. South Africa.

Araeotanypus pentheri Petrovitz, 1967. South Africa.

Araeotanypus striatus Schmidt, 1912. Tanzania.

Araeotanypus zumpti Petrovitz, 1967. South Africa.

 

Celaenochrous Kuijten, 1984 (1).

*Celaenochrous sinensis Kuijten, 1984. China.

 

Coilodes Westwood, 1845 (9).

Coelodes (incorrect subsequent spelling by Lacordaire (1856))

Coilodes castaneus Westwood, 1846. Colombia, Costa Rica, Nicaragua.

Coilodes chilensis Westwood, 1846. Chile? (doubtfull distribution).

Coilodes fumipennis Arrow, 1909. Brazil.

*Coilodes gibbus (Perty, 1830). (Hybosorus). Brazil.

Coilodes testaceus Pic, 1928. (synonym).

Hybosorus brasiliensis (Laporte, 1840). (synonym).

Coilodes humeralis (Mannerheim, 1829). (Hybosorus). Brazil.

            Hybosorus niger (Mannerheim, 1829). (synonym).

Hybosorus auger Westwood, 1846. (synonym).

Coilodes nigripennis Arrow, 1903. St. Vincent.

Coilodes ovalis Robinson, 1948. Venezuela.

Coilodes parvulus Westwood, 1846. Brazil.

Coilodes punctipennis Arrow, 1909. Ecuador, Peru.

 

 Coprologus Heer, 1847 (1).

*Coprologus gracilis Heer, 1847. Germany.

 

Cretohybosorus Nikolajev, 1999 (2).

*Cretohybosorus buryaticus Nikolajev, 1999. Russia (Siberia).

Cretohybosorus striatulus Nikolajev, 1999. Russia (Siberia).

 

Dicraeodon Erichson, 1847 (3).

Aporolaus Bates, 1887 (synonym).

*Dicraeodon basalis (Westwood, 1846). (Chaetodus? West.). French Guiana.

Dicraeodon fimbriatus (Bates, 1887). (Aporolaus). Panama.

Dicraeodon punctatus Arrow, 1911. Guatemala, Colombia.

 

Hapalonychoides Martínez, 1994 (1).

*Hapalonychoides similis Martínez, 1994. Argentina, Paraguay.

 

Hybosoroides Benderitter, 1914 (1).

*Hybosoroides alluaudi Benderitter, 1914. Kenya.

 

Hybosorus MacLeay, 1819 (5).

Hybosorus crassus Klug, 1855. Congo, Zimbabwe, Mozambique.

*Hybosorus illigeri Reiche, 1853. Africa, U.S.A., Mexico, Cuba, Haiti, Bahamas, Nicaragua, Venezuela, Israel, Saudi Arabia, Iran, Central Asia, Afghanistan, Pakistan, India, China, Vietnam, Portugal, Spain, France, Italy, Hungary, Greece, Bulgaria, Cyprus.

Scarabaeus  arator (Illiger, 1803) homonym of Scarabaeus arator (Fabricius 1792)

Hybosorus pinguis Westwood, 1845. (synonym).

Hybosorus roei Westwood, 1845. (synonym).

Hybosorus carolinus LeConte, 1847. (synonym).

Hybosorus nitidus Lansberge, 1882. (synonym).

Hybosorus illigeri nossibianus Fairmaire, 1895 (synonym).

Hybosorus. illigeri palearcticus Endrödi, 1957. (synonym).

Hybosorus laportei Westwood, 1845. Egypt, Ethiopia, Mauritania, Mali, Nigeria, Senegal, Togo.

            Hybosorus thoracicus Westwood, 1845. (synonym)

Hybosorus orientalis Westwood, 1845. Afghanistan, Myanmar, India, Indonesia (Java), Pakistan, Sri Lanka, Indonesia.

Hybosorus ruficornis Boheman, 1857. Botswana, Kenya, Mozambique, Namibia,Somalia, South Africa, Zambia, Zimbabwe.

 

 Hypseloderus Fairmaire, 1893 (1).

*Hypseloderus denticollis Fairmaire, 1893a. Vietnam.

 

Kuijtenous Paulian, 1981 (2).

*Kuijtenous laeviceps (Fairmaire, 1893). (Hybosorus). Comores Islands, Madagascar.

Hybosorus baliensis Brancsik, 1893 (synonym).

Phaeochrous insularis Linell, 1897 (synonym)

Kuijtenous tenuepunctatus (Fairmaire, 1895) (Hybosorus). Madagascar.

Hybosorus sparsepunctatus Pic, 1930 (synonym).

 

 Metachaetodus de Borre, 1886 (2).

Metachaetodus brunneicollis de Borre, 1886, Argentina.

*Metachaetodus discus de Borre, 1886, Uruguay.

 

Microphaeochroops Pic, 1930 (5).

Microphaechroops Pic 1930. (inadvertent error)

*Microphaeochroops hirsutus Pic, 1930. Vietnam.

Microphaeochroops laetus Arrow, 1942. Malaysia.

Microphaeochroops nigrosetosus Kuijten, 1985. Malaysia.

Microphaeochroops peninsularis Arrow, 1942. Malaysia.

Microphaeochroops varius Kuijten, 1985. Malaysia.

 

Microphaeolodes Kuijten, 1985 (1).

*Microphaeolodes mulumontis Kuijten, 1985. Malaysia.

 

Mimocoelodes Pic, 1930 (1).

*Mimocoelodes minutus Pic, 1930. Vietnam.

 

Pantolasius Lansberge, 1887 (2).

Pantolasius bandaharae Kuijten, 1985. Indonesia.

*Pantolasius vethi Lansberge, 1887. Indonesia.

 

Phaeochridius Lansberge, 1887 (2).

*Phaeochridius derasus (Harold, 1880). (Liparochrus).          Indonesia.

Phaeochridius haroldi Fairmaire, 1896. (synonym)

Phaeochridius uniformis Arrow, 1925. Malaysia (Sabah).

Phaeochridius benderitteri Pic, 1928. (synonym)

Phaeochridius cinereicollis Arrow, 1942. (synonym)

 

Phaeochroops CandŹze, 1876 (26).

Paeochroops (incorrect subsequent spelling by Nomura (1973)

Phaechroops (incorrect subsequent spelling by Pic (1928)

Phaeochroops acuticollis (Arrow, 1907) (Phaeochridius). Malaysia (Sabah).

Phaeochroops angulatus Benderitter, 1923. Philippines.

Phaeochroops colopacilis Kuijten, 1981. Vietnam.

Phaeochroops curtulus Schmidt, 1912. India.

Phaeochroops freenae Kuijten, 1981. Malaysia.

Phaeochroops gigas Arrow, 1907. Malaysia.

Phaeochroops gilleti Benderitter, 1923. Malaysia, Sabah, Philippines.

Phaeochroops punctulatus Arrow, 1942. (synonym).

Phaeochroops hisaeae Nishikawa, 1989. Malaysia, Sabah, Indonesia.

Phaeochroops indicus Arrow, 1907. India.

Phaeochroops lakhonicus Kuijten, 1981. Thailand.

*Phaeochroops lansbergei Candèze, 1876. Indonesia.

Phaeochroops laotianus Paulian, 1945. Laos.

Phaeochroops longisetosus Kuijten, 1981. India.

Phaeochroops maruyamai Nishikawa, 1989. Malaysia, Sabah, Indonesia.

Phaeochroops masumotoi Nishikawa, 1996. Thailand.

Phaeochroops meghalayicus Keith, 2001. India.

Phaeochroops myanmarensis Keith, 2000. Myanmar.

Phaeochroops ninbin Kuijten, 1981b. Vietnam.

Phaeochroops opacicollis Arrow, 1909. Myanmar, Thailand.

Phaeochroops peninsularis Arrow, 1909. Malaysia.

Phaeochroops rattus Arrow, 1909. Indonesia, Malaysia.

Phaeochroops batuensis Arrow, 1909. (synonym).

Phaeochroops mentaweiensis Arrow, 1909. (synonym).

Phaeochroops niasianus Arrow, 1909. (synonym).

Phaeochroops seres Kuijten, 1981. Vietnam.

Phaeochroops silphoides Fairmaire, 1898. Indonesia, Malaysia, Sabah, Philippines.

Phaeochroops recticollis Pic, 1928. (synonym).

Phaeochroops taiwanus Nomura, 1973. China (Taiwan).

Phaeochroops vulpecula Arrow, 1909. Indonesia, Malaysia.

Phaeochroops vulturius Kuijten, 1981. Indonesia.

 

Phaeochrous Laporte, 1840  (51).

Silphodes Westwood, 1845 (synonym).

Phaeochrous amplus Arrow, 1909. Cameroon.

Phaeochrous colmanti Schouteden, 1918 (synonym).

Phaeochrous bayeri Schouteden, 1918 (synonym).

Phaeochrous confusus Schouteden, 1918 (synonym).

Phaeochrous westwoodi Schouteden, 1918 (synonym).

Phaeochrous australicus Kuijten, 1978. Australia.

Phaeochrous beccarii Harold, 1871. Ethiopia, Sudan, Tanzania, Yemen.

Phaeochrous arabicus Arrow, 1909 (synonym).

Phaeochrous elgonensis Schouteden, 1918 (synonym).

Phaeochrous gonsalvesi Petrovitz, 1975 (synonym).

Phaeochrous niloticus Burgeon, 1928 (synonym).

Phaeochrous stercorarius Kolbe, 1895 (synonym).

Phaeochrous tangensis Schouteden, 1918 (synonym).

Phaeochrous becarii, Schmidt, 1913 (synonym).

Phaeochrous bicarinatus Kuijten, 1986. Uganda, Rwanda, Burundi

Phaeochrous borealis Kuijten, 1984. China.

Phaeochrous burgoblitus Kuijten, 1986. Zaire, Uganda, Rwanda.

Phaeochrous kivuensis Burgeon. (nomen nudum).

Phaeochrous camerunensis Arrow, 1909. Cameroon, Gabon, Nigeria, Congo, Zaire, Sudan, Ethiopia, Tanzania, Zambia, Rwanda, Burundi, Uganda, Mozambique, South Africa.

Phaeochrous boranus Müller, 1939 (synonym).

Phaeochrous mashunus Arrow 1909 (synonym).

Phaeochrous zombensis Schouteden, 1918 (synonym).

Phaeochrous vicinus Schouteden, 1918 (synonym).

Phaeochrous compactus Kuijten, 1978. Sri Lanka.

Phaeochrous davaonis Kuijten, 1981. Philippines.

Phaeochrous dispar Quedenfeldt, 1884. Angola, Zaire.

Phaeochrous thomensis Arrow, 1909 (synonym).

Phaeochrous kuiluensis Schouteden, 1918 (synonym).

Phaeochrous dissimilis dissimilis Arrow, 1909. Myanmar, China, Laos, Thailand, Vietnam.

Phaeochrous dissimilis vietnamicola Kuijten, 1978. China, Vietnam.

Phaeochrous diversipes Pic, 1928. India.

Phaeochrous dubius (Westwood, 1846). (Silphodes). Indonesia.

Phaeochrous elevatus Kuijten, 1978. Sri Lanka.

Phaeochrous emarginatus benderitteri Pic, 1928. Indonesia (Waigeu Island).

Phaeochrous rugosicollis Benderitter, 1913. (synonym).

Phaeochrous celebensis var. ruficeps Pic. 1928. (synonym).

Phaeochrous emarginatus buruensis Kuijten, 1978. Indonesia (Buru Island)

Phaeochrous emarginatus davidis Fairmaire, 1886. China.

*Phaeochrous emarginatus emarginatus Laporte, 1840. Japan, Melanesia, China, India, Thailand, Buthan, Laos, Vietnam, Malaysia, Indonsesia, Philipines, Papua New Guinea, Australia.

Phaeochrous sumatrensis (Westwood, 1846) (Silphodes). (synonym).

Phaeochrous hirtipes (MacLeay, 1864) (Silphodes). (synonym).

Phaeochrous alternatus Fairmaire, 1879 (synonym).

Phaeochrous asiaticus Lewis, 1896 (synonym).

Phaeochrous pallidus Arrow, 1909 (synonym).

Phaeochrous celebensis Pic, 1928 (synonym).

Phaeochrous perroudi Pic, 1943 (synonym).

Phaeochrous gracilis Petrovitz, 1975 (synonym).

Phaeochrous emarginatus thilliezi Keith, 1999. Pakistan.

Phaeochrous emarginatus suturalis Lansberge, 1885. (Myanmar).

Phaeochrous enigmaticus Kuijten, 1978. Bangladesh, China, Indonesia.

Phaeochrous gambiensis (Westwood, 1846) (Silphodes). Gambia, Senegal, Burkina, Chad.

Phaeochrous gigas Schouteden, 1918. Africa.

Phaeochrous latus Pic, 1944 (synonym).

Phaeochrous hainanensis Zhang-Youwei, 1990. China.

Phaeochrous indicus (Westwood, 1846). (Silphodes). India.

Phaeochrous intermedius intermedius Pic, 1928. Bhutan, China, India, Laos, Thailand, Vietnam.

Phaeochrous intermedius occidentalis Kuijten, 1978. Bhutan.

Phaeochrous lobatus Kuijten, 1978. Philippines.

Phaeochrous madagascariensis (Westwood, 1846). (Silphodes). Madagascar.

Phaeochrous madrassicus Kuijten, 1978. India.

Phaeochrous nicolasi Keith, 2002. Australia.

Phaeochrous nitidus Arrow, 1909. Tanzania.

Phaeochrous arrowi Schouteden 1918 (synonym).

Phaeochrous philippinensis (Westwood, 1846) (Silphodes). Philippines.

Phaeochrous pletus Kuijten, 1978. East Timor.

Phaeochrous portuum Kuijten, 1978. New Guinea.

Phaeochrous pseudintermedius Kuijten, 1978. Vietnam.

Phaeochrous rhodesianus Schouteden, 1918. Zimbabwe.

Phaeochrous kapiriensis Schouteden, 1918 (synonym).

Phaeochrous rudis Kuijten, 1984. Indonesia.

Phaeochrous ruficollis Fairmaire, 1893a. Vietnam.

Phaeochrous rufus Pic, 1928b. China.

Phaeochrous schoutedeni Burgeon, 1928. Central Africa.

Phaeochrous separabilis Zhang-Youwei, 1990. China.

Phaeochrous stupendus Kuijten, 1986. Equatorial Africa.

Phaeochrous sulawesi Kuijten, 1978. Indonesia.

Phaeochrous tanzanicus Tagliaferri, 2002. Tanzania.

Phaeochrous tonkineus Pic, 1943. Vietnam.

Phaeochrous tumbanus Burgeon, 1928. Angola, Ivory Coast, Congo, Zambia, Central African Republic, Gabon, Ethiopia, Gabon, Kenya, Rwanda, Burundi, Tanzania, Uganda.

Phaeochrous tokaraensis Nomura, 1961. Japan.

Phaeochrous turcanicola Kuijten, 1986. Kenya.

Phaeochrous uelensis Burgeon, 1928. Gabon, Cameroon, Zaire, Uganda, Mozambique, Madagascar.

Phaeochrous usambarae Burgeon, 1928. Tanzania.

 

Phaeocroides Péringuey, 1908 (5).

Phaeochroides (incorrect subsequent spelling by Schmidt (1913).

*Phaeocroides damarinus Péringuey, 1908. Namibia.

Phaeocroides effetus (Kolbe, 1907). (Phaeocrhous?). Namibia, Botswana.

            Phaeocroides juvenilis Kolbe, 1912. (synonym).

Phaeocroides nanus (Arrow, 1942). (Phaeocrhous). India, Pakistan.

Phaeocroides mapellii Petrovitz, 1972. Bangladesh.

Phaeocroides orientalis (Petrovitz, 1963). (Araeotanopus).   India.

 

Procoilodes Ocampo, 2002 (1).

*Procoilodes adrastus Ocampo, 2002. Dominican Republic.

 

Seleucosorus Kuijten, 1983 (1).

*Seleucosorus punctatissumus (Reiche, 1861) (Hybosorus). Syria, Turkey.

 

Tyrannasorus Ratcliffe and Ocampo, 2001 (1).

*Tyrannasorus rex Ratcliffe and Ocampo, 2001. Dominican Republic.

 

Liparochrinae Ocampo

 Type species Liparochrous Erichson, 1848.

 Antiochrus Sharp, 1873 (7).

Antiochrus aberrans (Fairmaire, 1877). (Liparaochrus). Australia.

*Antiochrus brunneus Sharp, 1873. Australia.

Antiochrus oblongus Harold, 1873 (synonym).

Antiochrus freyi Petrovitz, 1963. Australia.

Antiochrus politulus (MacLeay, 1888). (Liparaochrus). Australia.

Antiochrus septentrionalis Petrovitz, 1968. Australia.

Antiochrus setosus Petrovitz, 1963. Australia.

Antiochrus similis Petrovitz, 1971. Australia.

   

Liparochrus Erichson, 1848 (41).

Ranidia Paulian, 1980 (synonym).

Subgenus uncertain.

Liparochrus asperulus Fairmaire, 1877. Australia.

Subgenus Liparochrus Erichson 1848.

Liparochrus (Liparochrus) bimaculatus Macleay, 1864. Australia.

Liparochrus (Liparochrus) crassicollis Arrow, 1925. Papua New Guinea.

Liparochrus (Liparochrus) dilatatifrons Blackburn, 1905. Australia.

Liparochrus dilatifrons Arrow, 1912. (misspelling).(synonym).

Liparochrus timidus Arrow, 1909. (synonym).

Liparochrus (Liparochrus) dolium Heller, 1912. Papua New Guinea.

Liparochrus (Liparochrus) dux Arrow, 1909. Papua New Guinea.

Liparochrus (Liparochrus) eungellae Paulian, 1980. Australia.

Liparochrus (Liparochrus) fossulatus Westwood, 1852. Australia.

*Liparochrus (Liparochrus) geminatus Westwood, 1852. Australia.

Liparochrus (Liparochrus) hackeri Blackburn, 1912. Australia.

Liparochrus freyi Petrovitz, 1963. (synonym).

Liparochrus (Liparochrus) insularis Petrovitz, 1980, Lord Howe Island.

Liparochrus (Liparochrus) krikkeni Paulian, 1980. Papua New Guinea.

Liparochrus (Liparochrus) laevipennis Petrovitz, 1963. Australia.

Liparochrus (Liparochrus) laevis Paulian, 1980. Australia.

Liparochrus (Liparochrus) laevissimus Paulian, 1980a. Australia.

Liparochrus (Liparochrus) matthewsi Paulian, 1980a. New Caledonia.

Liparochrus (Liparochrus) nanus Paulian, 1980a. Australia.

Liparochrus (Liparochrus) nitidicollis Blackburn, 1905. Australia.

Liparochrus (Liparochrus) occidentalis Paulian, 1980. Australia.

Liparochrus (Liparochrus) papuus Lansberge, 1885. Papua New Guinea.

Liparochrus alternatus Macleay, 1886 (synonym).

Liparochrus (Liparochrus) quadrimaculatus Harold, 1877. Australia.

Liparochrus (Liparochrus) rufus Blackburn, 1892. Australia.

Liparochrus (Liparochrus) sculptilis Westwood, 1852. Australia.

Liparochrus ciliboides Harold, 1875 (synonym).

Liparochrus (Liparochrus) septentrionalis Paulian, 1980. Australia.

Liparochrus (Liparochrus) silphoides Harold, 1874. Australia.

Liparochrus raucus Fairmaire, 1877 (synonym).

Liparochrus (Liparochrus) sulcatus (Montrouzier, 1860). (Sphaeridium). New Caledonia.

                        Sphaeridium sulcatus Montrouzier, 1860 (synonym).

Subgenus Parolichrus Allsopp, 1982.

*Liparochrus (Parolichrus) ingens Felsch, 1909. Papua New Guinea.

Liparochrus ingenti, Heller 1912. (synonym).

Liparochrus (Parolichrus) lugubris Arrow, 1925. Indonesia, Papua New Guinea.

Subgenus Ranidichruss Allsopp, 1982.

Liparochrus (Ranidichrus) cahilli Paulian, 1980. Australia.

Liparochrus (Ranidichrus) carnei Paulian, 1980. Australia.

Liparochrus (Ranidichrus) crenatulus Fairmaire, 1877. Australia.

Liparochrus crenulatus, Lansberge, 1885. (synonym).

Liparochrus weyersi Petrovitz, 1971. (synonym).

Liparochrus (Ranidichrus) darlingtoni Paulian, 1980. Australia.

Liparochrus (Ranidichrus) demarzi Petrovitz, 1963. Australia.

Liparochrus (Ranidichrus) globuliformis Macleay, 1888. Australia.

Liparochrus (Ranidichrus) infantus Petrovitz, 1963. Australia.

Liparochrus (Ranidichrus) irianae Paulian, 1980. Indonesia, Papua New Guinea.

Liparochrus (Ranidichrus) modestus Paulian, 1980. Papua New Guinea.

*Liparochrus (Ranidichrus) multistriatus Harold, 1874. Australia.

Liparochrus pimelioides Lansberge, 1885. (synonym).

Liparochrus pimeloides, Schimdt, 1913. (misspelling).

Liparochrus (Ranidichrus) septemdecimlineatus Petrovitz, 1968. Australia.

Liparochrus (Ranidichrus) tropicus Petrovitz, 1963. Australia.

Subgenus Ropalichrus Allsopp, 1982.

Liparochrus (Ropalichrus) monteithi Paulian, 1980. Australia.

*Liparochrus (Ropalichrus) storeyi Paulian, 1980. Australia.

 

Pachyplectrinae Ocampo 

Type species Pachyplectrus LeConte, 1874. 

Pachyplectrus LeConte, 1874 (1).

*Pachyplectrus laevis LeConte, 1874. U.S.A.

 

Brenskea Reitter, 1891 (2).

Spermohybosorus Pic 1922. (synonym).

            Brenskea chudeaui Reitter, 1909. Algeria.

*Brenskea coronata Reitter, 1891. Central Asia, Iran, Israel, Morocco, Algeria, Tunisia, Libya,

Egypt, Azerbaijan, Russia.

Brenskea varentzovi Semenov, 1896 (synonym)

                        Spermohybosorus testaceous Pic, 1922 (synonym).

 

Incertae sedis

Borrochrus Allsopp, 1979 (2).

Trichops de Borre, 1886. (preoccupied homonym). (type species Trichops ciliatus de Borre, nec Val, 1853).

*Borrochrus ciliatus (de Borre, 1886). (Trichops). Brazil, Paraguay, Argentina.

Borrochrus mutilus (Petrovitz, 1968). (Trichops). Paraguay.

 

 Daimothoracodes Petrovitz, 1970 (4).

Daimothoracodes confossus Ocampo and Vaz-de-Mello, 2002. Brazil.

Daimothoracodes magnificus Martínez, 1994. Bolivia, Argentina.

*Daimothoracodes mirabilis Petrovitz, 1970. Brazil.

Daimothoracodes rugomarginatus Ocampo, 2005. Brazil.