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CHECKLIST OF THE DYNASTINAE OF
HONDURAS, NICARAGUA, AND EL SALVADOR

Tribe Cyclocephalini
Ancognatha sellata Arrow, 1911

Aspidolea fuliginea Burmeister, 1847
Aspidolea singularis Bates, 1888
       Aspidolea texana Höhne, 1922 (synonym)
       Aspidolea similis Höhne, 1922 (synonym)
       Aspidolea cevallosi Martínez, 1975 (synonym)

Cyclocephala abrelata Ratcliffe and Cave, 2002
Cyclocephala alexi Ratcliffe and Delgado, 1992
Cyclocephala amblyopsis Bates, 1888
       Cyclocephala amblyopsis monochroa Bates, 1888 (synonym)
Cyclocephala ampliata Bates 1888
Cyclocephala atripes Bates, 1888
Cyclocephala batesi Delgado and Castañeda, 1994
Cyclocephala brevis Höhne, 1923
       Cyclocephala pubescens Burmeister, 1847 (primary junior homonym)
Cyclocephala carbonaria Arrow, 1911
       Mononidia trachypyga Prell, 1934 (synonym)
       Mononidia punctulata Prell, 1934 (synonym)
       Cyclocephala howdeni Endrödi, 1967 (synonym)
Cyclocephala complanata Burmeister, 1847
       Cyclocephala obliquata Casey, 1915 (synonym)
       Cyclocephala emacerata Casey, 1915 (synonym)
Cyclocephala concolor Burmeister, 1847
Cyclocephala confusa Endrödi, 1966
Cyclocephala conspicua Sharp, 1877
       Cyclocephala conspicua gregaroides Dechambre, 1992 (synonym)
       Cyclocephala conspicua fusca Dechambre, 1992 (synonym)
Cyclocephala curta Bates, 1888
       Cyclocephala fusciventris Arrow, 1902 (synonym)
Cyclocephala deceptor Casey, 1915
Cyclocephala discolor (Herbst, 1792)
       Melolontha unciata Schönherr, 1817 (synonym)
       Cyclocephala aurantiaca Prell, 1937 (synonym)
       Cyclocephala discolor andina Bréthes, 1904 (synonym)
Cyclocephala erotylina Arrow, 1914
Cyclocephala freudi Endrödi, 1963
Cyclocephala fulgurata Burmeister, 1847
Cyclocephala gravis Bates, 1888
Cyclocephala guttata Bates, 1888
Cyclocephala kaszabi Endrödi, 1964
Cyclocephala larssoni Endrödi, 1964
Cyclocephala ligyrina Bates, 1888
Cyclocephala lunulata Burmeister, 1847
       Cyclocephala nubeculosa Burmeister, 1847 (synonym)
       Graphalia oblita Casey, 1915 (synonym)
Cyclocephala maculiventris Höhne, 1923
Cyclocephala mafaffa Burmeister, 1847
       Cyclocephala grandis Burmeister, 1847 (synonym)
       Stigmalia cuernavacana Casey, 1915 (synonym)
       Stigmalia fallaciosa Casey, 1915 (synonym)
       Stigmalia deficiens Casey, 1915 (synonym)
       Stigmalia mafaffa histrionica Casey, 1915 (synonym)
Cyclocephala melanocephala (Fabricius, 1775)
       Melolontha leucophthalma Fischer von Waldheim, 1823 (synonym)
       Melolontha ventralis Erichson, 1847 (synonym)
       Cyclocephala dimidiata Burmeister, 1847 (synonym)
       Cyclocephala elegans Horn, 1871 (synonym)
       Dichromina ocularis Casey, 1915 (synonym)
Cyclocephala melolonthida Ratcliffe and Cave, 2002
Cyclocephala multiplex Casey, 1915
Cyclocephala mutata Harold, 1869
       Cyclocephala laevicauda Arrow, 1902 (synonym)
       Cyclocephala pseudisabellina Endrödi, 1980 (synonym)
       Cyclocephala vitracelis Dechambre, 1999 (synonym)
Cyclocephala ovulum Bates, 1888
Cyclocephala picopijola Ratcliffe and Cave, 2006
Cyclocephala porioni Dechambre, 1979
Cyclocephala prolongata Arrow, 1902
Cyclocephala sanguinicollis Burmeister, 1847
       Cyclocephala divisa Casey, 1915 (synonym)
       Cycocephala politicauda Casey, 1915 (synonym)
Cyclocephala sexpunctata Laporte, 1840
       Cyclocephala pubescens Erichson, 1847 (synonym)
       Cyclocephala lucida Burmeister, 1847 (synonym)
       Cyclocephala sexpunctata spermophila Ohaus, 1910 (synonym)
       Stigmalia triangulifer Casey, 1915 (synonym)
       Stigmalia discoidalis Casey, 1915 (synonym)
       Stigmalia costaricana Casey, 1915 (synonym)
       Stigmalia circulifer Casey, 1915 (synonym)
       Cyclocephala pubescens nigripes Höhne 1923 (synonym)
Cyclocephala sororia Bates, 1888
       Cyclocephala flava Dechambre, 1999 (synonym)
Cyclocephala sparsa Arrow, 1902
       Cyclocephala landini Endrödi, 1964 (synonym)
       Cyclocephala virkii Howden and Endrödi, 1966 (synonym)
Cyclocephala stictica Burmeister, 1847
       Cyclocephala sexnotata Burmeister, 1847 (synonym)
       Cyclocephala microspila Bates, 1888 (synonym)
       Cyclocephala stictica bilineata Höhne, 1923 (synonym)
Cyclocephala tutilina Burmeister, 1847
Cyclocephala weidneri Endrödi, 1964

Dyscinetus dubius (Olivier, 1789)
       Melolontha geminatus Fabricius, 1801 (synonym)
       Geotrupes lugubris Quensel in Schönherr, 1806 (synonym)
       Dyscinetus frater Bates, 1888 (synonym)
Dyscinetus obtusus Casey, 1915 (synonym)
Dyscinetus laevipunctatus Bates, 1888
       Palechus histrio Casey, 1915 (synonym)

Erioscelis columbica Endrodi, 1966
Mimeoma acuta Arrow, 1902

Stenocrates bicarinatus Robinson, 1947
       Stenocrates difficilis Endrödi, 1966 (synonym)
Stenocrates canuli Delgado, 1991
Stenocrates hardyi Dechambre, 1985
Stenocrates laevicollis Kirsch, 1871

Tribe Pentodontini

Bothynus complanus (Burmeister, 1847)
       Bothynus simplicitarsis Bates, 1888 (synonym)
       Bothynus monstrosus Bates, 1888 (synonym)
Bothynus quadridens (Taschenberg, 1870)

Euetheola bidentata (Burmeister, 1847)
       Cyclocephala brevis Perty, 1830 (synonym?  Type unknown)
       Heteronychus globosus Burmeister, 1847 (synonym?  Type unknown)
Euetheola humilis (Burmeister, 1847)
       Ligyrus rugiceps LeConte, 1856 (synonym)
       Dyscinetus parvus Casey, 1915 (synonym)
       Dyscinetus hondurana Casey, 1915 (synonym)

Orizabus botox Ratcliffe and Cave,2006
Orizabus clunalis (LeConte, 1856)
Orizabus hondurensis Ratcliffe and Cave, 2006
Orizabus puchicus Ratcliffe and Cave, 2006
Orizabus teamscaraborum Ratcliffe and Cave,2006

Parapucaya amazonica Prell, 1934

Tomarus bituberculatus (Palisot de Beauvois, 1811)
       Ligyrus latifovea Bates, 1888 (synonym)
       Ligyrus maximus Arrow, 1913 (synonym)
       Ligyrus latus Arrow, 1914 (synonym)
Tomarus cicatricosus (Prell, 1937)
Tomarus ebenus (Degeer, 1774)
       Scarabaeus cordatus Fabricius, 1792 (synonym)
       Cyclocephala scarabaeinus Perty, 1830 (synonym)
Tomarus fossor (Latreille, 1813)
       Ligyrus castaneipennis Apolinar, 1927 (synonym)
Tomarus gyas Erichson, 1848
       Ligyrus amazonicus Arrow, 1914 (synonym)
Tomarus laevicollis (Bates, 1888)
       Ligyrus bryanti Rivers, 1891 (synonym)
Tomarus maternus (Prell, 1937)
Tomarus nasutus (Burmeister, 1847)
       Ligyrus pygidialis Bates, 1888 (synonym)
Tomarus sallaei (Bates, 1888)
       Ligyrodes propinquus Casey, 1915 (synonym)
       Ligyrodes aztecus Casey, 1915 (synonym)

Tribe Oryctini

Coelosis biloba (Linnaeus, 1767)
       Coelosis biloba lepesmei Bourgin, 1944  (invalid name)
       Coelosis biloba incana Bourgin, 1944  (invalid name)
       Coelosis biloba cacica Bourgin, 1944  (invalid name)
       Coelosis biloba tibialis Bourgin, 1944 (synonym)
       Coelosis biloba tibialis pauliani Bourgin, 1944 (invalid name)

Enema endymion Chevrolat, 1843
       Enema lupercus Burmeister, 1847 (synonym)
       Enema paniscus Burmeister, 1847 (synonym)
       Enema gibbicollis Sternberg, 1908 (synonym)
Enema pan (Fabricius, 1775)
       Scarabaeus chorinaeus Fabricius, 1775 (synonym)
       Scarabaeus quadrispinosus Fabricius, 1781 (synonym)
       Scarabaeus enema Fabricius, 1787 (synonym)
       Scarabaeus aeneas Kirby, 1818 (synonym)
       Scarabaeus titornus Perty, 1830 (synonym)
       Enema lupercus Chevrolat, 1843 (synonym)
       Enema infundibulum Burmeister, 1847 (synonym)

Heterogomphus chevrolati Burmeister, 1847
       Heterogomphus chevrolati eurytus Bates, 1888 (synonym)
       Heterogomphus chevrolati punctatissimus Prell, 1912 (synonym)
       Heterogomphus chevrolati insignis Prell, 1912 (synonym).
Heterogomphus mniszechi (Thomson, 1859)
       Daemonoplus pehlkei Kolbe 1906a: 273 (synonym)
       Daemonoplus kolbei Prell 1912b: 155 (synonym)

Irazua dilicra Ratcliffe, 2003

Megaceras morpheus Burmeister, 1847
       Megaceras ixyon Reiche, 1859 (synonym)
Megaceras septentrionis Bates, 1888
       Megaceras septentrionis crassicornis Dechambre, 1975 (synonym)

Podischnus agenor (Olivier, 1789)
       Scarabaeus barbicornis Latreille, 1812 (synonym)
       Podischnus propinquus Prell, 1911 (synonym)

Strategus aloeus (Linnaeus, 1758).
       Geotrupes semiramis Fabricius, 1801 (synonym)
       Scarabaeus aesalus Laporte, 1840 (synonym)
       Strategus julianus Burmeister, 1847 (synonym)
       Strategus piosomus Kolbe, 1906 (synonym)
       Strategus roosevelti Casey, 1915 (synonym)
       Strategus frontalis Casey, 1915 (synonym)
       Strategus tarsalis Casey, 1915 (synonym)
       Strategus gaillardi Casey, 1915 (synonym)
Strategus hipposiderus Ratcliffe, 1976
Strategus jugurtha Burmeister, 1847
Strategus longichomperus Ratcliffe, 1976

Xyloryctes ensifer Bates, 1888
Xyloryctes lobicollis Bates, 1888
Xyloryctes splendidus Prell, 1914
Xyloryctes teuthras Bates, 1888

Tribe Phileurini

Amblyodus taurus Westwood, 1878
Amblyoproctus chalumeaui Endrödi, 1977

Archophileurus simplex (Bates, 1888)
Goniophileurus femoratus (Burmeister, 1847)

Hemiphileurus cavei Ratcliffe, 2003
Hemiphileurus cylindroides (Bates, 1888)
       Hemiphileurus costaricensis Endrödi, 1978 (synonym)
       Hemiphileurus jamesonae Ratcliffe, 1988 (synonym)
Hemiphileurus dejeani (Bates, 1888)
Hemiphileurus euniceae Ratcliffe and Cave, 2006
Hemiphileurus laevicauda (Bates, 1888)
Hemiphileurus simplex  (Prell, 1914)
       Hemiphileurus fraternus Arrow, 1937 (synonym)
Hemiphileurus variolosus (Burmeister, 1847)
       Epiphileurus irregularis Prell, 1914 (synonym)
       Hemiphileurus variolosus striatus Endrödi, 1978 (synonym)

Homophileurus luedeckei Kolbe, 1910
Homophileurus quadrituberculatus (Palisot de Beauvois, 1806)
       Scarabaeus bajulus Perty, 1830 (synonym)
       Phileurus cephalotes Laporte, 1840 (synonym)
       Homophileurus muticus Prell, 1914 (synonym)
Homophileurus tricuspis Prell, 1914

Phileurus carinatus Prell, 1914
       Phileurus carinatus declivis Prell, 1914 (synonym)
Phileurus didymus (Linnaeus, 1758)
       Phileurus affinis Reiche, 1859 (synonym)
Phileurus truncatus Palisot de Beauvois, 1807
       Phileurus recurvatus Casey, 1915 (synonym)
Phileurus valgus (Olivier, 1789)
       Scarabaeus castaneus Haldeman, 1843 (synonym)
       Phileurus capra Bates, 1888 (synonym)
       Phileurus valgus septentrionis Kolbe, 1910 (synonym)
       Phileurus meridionalis Kolbe, 1910 (synonym)
       Phileurus valgus antillarum Prell, 1912 (subspecies)
       Phileurus texensis Casey, 1915 (synonym)
       Phileurus carolinae Casey, 1915 (synonym)
       Phileurus sulcifer Casey, 1915 (synonym)
       Phileurus floridanus Casey, 1915 (synonym)
       Phileurus clathratus Casey, 1915 (synonym)
Phileurus voirinae Endrödi, 1985
Phileurus youngi Ratcliffe, 1988

Tribe Agaocephalini

Spodistes beltianus (Bates, 1888)
       Lycomedes reichii Burmeister, 1847 (synonym)
Spodistes mniszechi (Thomson, 1860)
Spodistes monzoni Warner, 1992

Tribe Dynastini

Dynastes hercules (Linnaeus, 1758)
            Scarabaeus scaber Linnaeus, 1764 (synonym)
            Scarabaeus hercules oculatus Scopoli, 1772 (synonym)
            Scarabaeus alcides Fabricius, 1781 (synonym)
            Scarabaeus iphiclus Panzer, 1782 (synonym)
            Scarabaeus perseus Olivier, 1789 (synonym)
            Dynastes lagaii Verrill, 1906 (synonym)
            Dynastes vulcan Verrill, 1906 (synonym)
            Dynastes argentatus Verrill, 1907 (synonym)
            Dynastes hercules ecuatorianus Ohaus, 1913 (synonym)
            Dynastes hercules niger Endrödi, 1947 (synonym)
            Dynastes hercules baudrii Pinchon, 1976 (synonym)
            Dynastes hercules reidi Chalumeau, 1977 (synonym)
            Dynastes hercules septentrionalis Lachaume, 1985 (synonym)
            Dynastes hercules occidentalis Lachaume, 1985 (synonym)
            Dynastes hercules lichyi Lachaume, 1985 (synonym)
            Dynastes hercules paschoali Grossi and Arnaud, 1993 (synonym)
            Dynastes hercules tuxtlaensis Morón, 1993 (synonym)
            Dynastes hercules bleuzeni Silvestre and Dechambre, 1995 (synonym)
            Dynastes hercules  trinidadensis Chalumeau and Reid, 1995 (synonym)
            Dynastes hercules morishimai Nagai, 2002 (synonym)
            Dynastes hercules takakuwai Nagai, 2002 (synonym)
Dynastes maya Hardy, 2003

Golofa pizarro Hope, 1837
       Golofa hastata Burmeister, 1847 (synonym)
       Golofa imperialis Thomson, 1858 (synonym)
       Golofa sallei Thomson, 1860 (synonym)
       Golofa clavicornis Thomson, 1860 (synonym)
Golofa tersander Burmeister, 1847
       Mixigenus leander Thomson, 1859 (synonym)
       Mixigenus barbicornis Fairmaire, 1878 (synonym)
       Golofa dohrni Nonfried 1890, (synonym)

Megasoma elephas (Fabricius, 1775)
       Megasoma mexicanum Fischer, 1968 (synonym)
       Megasoma elephas iijimai Nagai, 2003 (synonym)

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
   
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