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        Psammodius Fallén 1807 
        Psammodius Fallén 1807: 37. 
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            Psammodius basalis (Mulsant and Rey) 
            Photo by Paul Skelley.  | 
           
         
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      Type species. Scarabaeus asper Fabricius 1775: 20, by subsequent designation (= Aphodius sulcicollis Illiger 1801: 20).                                     
              = Psammobius Heer 1841: 531.Type species: Aphodius  
               sulcicollis Illiger 1801, by subsequent designation. 
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    Diagnosis. Body length 3.0-4.0 mm, elongate to weakly globose, reddish-brown to almost black, dorsal surfaces glabrous, with lateral fringe of setae at least on pronotum. Head surface coarsely granulate, frontal suture indistinct, vertex with 1-2 pair V-shaped ridges. Clypeus rounded each side of median emargination. Pronotum with distinct set of 4 transverse grooves and 5 ridges. Elytral base finely margined; intervals moderately convex. Metafemur robust. Metatibia weakly curved, widened at apex. Metatarsi shortened; basal metatarsomere triangular.  | 
  
  
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    Distribution. Africa, Palearctic to Oriental region, northeastern North America (introduced).  | 
  
  
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    Composition. Worldwide, Psammodius contains 20 species, 3 of which are introduced in the New World.  
           
          Psammodius basalis (Mulsant and Rey 1869: 636) [Psammobius], Europe, NE-USA (introduced). 
          Psammodius laevipennis Costa 1844: 18, Europe, NE-USA (introduced). 
                  = Psammodius aegialioides Haldeman 1848: 107. 
                  = Psammodius ciliatus Kuster 1849: 51.  
                  = Psammodius rugicollis Erichson 1848: 916. 
                  = Psammodius scutellaris Mulsant and Wachanru 1859: 187.  
Psammodius pierotti Pittino 1979: 598, Europe, western Asia, NE-USA (introduced). 
                   = Aphodius sulcicollis Illiger 1801: 20, of authors [not  
                   Psammodius asper (Fab. 1775)].  | 
  
  
  
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    Life History. In North America, Psammodius were possibly introduced in ships ballast. Kim and Lumaret (1981) discuss the biology for the related P. porcicollis (Illiger) in Europe.  | 
  
  
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    Larvae. The larvae of P. pierotti was described by Medvedev (1952).  | 
  
  
  
  
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      References: 
      Cartwright, O. L. 1955. Scarab beetles of the genus Psammodius in the Western Hemisphere. Proceedings of the United States National Museum 104(3344): 413-462.  
            Kim, J. I., and J. P. Lumaret. 1981. Ecologie de Psammodius porcicollis (Ill.) (Col., Aphodiidae), insecte sabulicole des dunes littorales du Golfe du Lion. Annales de la Societe Entomologique de France 17: 449‑462.  
        Medvedev, S. I. 1952. Larvae of scarabaeid beetles of the fauna of the U.S.S.R. Opredeliteli po Fauna U.S.S.R. Moscow 47: 1-242 
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            Author: Paul Skelley 
  
              Generated on: 09/MAY/08.....Last 
              modified: 09/MAY/08 
              University of Nebraska State Museum - Division of Entomology              | 
         
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